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keys to the vascular plants of east texas - Botanical Research ...

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RHYNCHOSPORA/CYPERACEAE<br />

621<br />

Lipocarpha aristulata (Coville) G.C. Tucker, (bearded or awned), AWNED HALFCHAFF SEDGE.<br />

Spikelet-like spikes 1–5(–8) mm long; floral scales widest near middle; inner hyaline scale 0.5–<br />

0.8 mm long. Disturbed wet soils, exposed shorelines, usually sandy soils; Comal and Travis<br />

(Turner et al. 2003) cos. near sw margin <strong>of</strong> East TX; also range map given by Friedland (1941)<br />

clearly showed this taxon within East TX (counties not specified); scattered in w 2/3 <strong>of</strong> TX; w<br />

U.S. from MI, MO, and TX w <strong>to</strong> <strong>the</strong> Pacific coast. Jun–Nov. [Hemicarpha aristulata (Coville)<br />

Smyth, Hemicarpha micrantha (Vahl) Pax var. aristulata Coville]<br />

Lipocarpha drummondii (Nees) G.C. Tucker, (for its discoverer, Thomas Drummond, 1780–1835,<br />

Scottish botanist and collec<strong>to</strong>r in North America), DRUMMOND’S HALFCHAFF SEDGE, COMMON<br />

HEMICARPHA. Spikelet-like spikes 1–5(–6) mm long; floral scales widest beyond middle. Disturbed<br />

wet soils, exposed shorelines, usually sandy soils; Smith (BRIT), Bastrop, Harrison, Lee,<br />

Leon, Travis, and Tyler (Turner et al. 2003) cos.; also Gulf Prairies and Marshes and e Cross Timbers<br />

and Prairies; c U.S. from OH s <strong>to</strong> TX w <strong>to</strong> NE and AZ. May–Nov. [Hemicarpha drummondii<br />

Nees, H. micrantha (Vahl) Pax var. drummondii (Nees) Friedl.]<br />

Lipocarpha micrantha (Vahl) G.C. Tucker, (small-flowered), SMALL-FLOWER HALFCHAFF SEDGE.<br />

Spikelet-like spikes 1–3(–5) mm long; floral scales widest at or beyond middle. Moist sandy<br />

soils, seasonally wet areas, exposed shorelines; Brazos, Harrison, Henderson (BRIT), and Lee<br />

(Turner et al. 2003) cos.; also Gulf Prairies and Marshes (Calhoun Co.—BRIT), Cross Timbers<br />

and Prairies (Bosque, Hood, and Parker (BRIT) cos.) and Trans-Pecos; s Canada and widespread<br />

in <strong>the</strong> U.S. except <strong>the</strong> n Rocky Mt. area. Mar–Oct. [Hemicarpha micrantha (Vahl) Pax,<br />

Hemicarpha micrantha var. minor (Schrad.) Friedl., Scirpus micranthus Vahl]<br />

RHYNCHOSPORA Vahl<br />

BEAK-RUSH, HORNED-RUSH, BEAK SEDGE, WHITE-TOP<br />

Tufted or clump-forming, sometimes rhizoma<strong>to</strong>us perennials (rarely annuals), glabrous or with<br />

scabrous-margined leaf blades; culms (= stems) triangular in cross section; leaves basal and<br />

cauline; spikelets narrowly ovoid <strong>to</strong> fusiform or roundish, usually brownish or white (in a few<br />

species) or dark brown <strong>to</strong> nearly black (in a few species), usually 1–few-flowered, in loose or<br />

compact clusters, in spike-like or open panicles, or in a head-like cluster, <strong>the</strong> inflorescence with<br />

1–few leafy bracts; scales <strong>of</strong> spikelets spirally arranged, <strong>the</strong> lower (1–)2–3 usually sterile; uppermost<br />

1–2 florets usually without pistil; perianth <strong>of</strong> bristles or absent; achenes flattened <strong>to</strong> lenticular<br />

<strong>to</strong> nearly round in cross section, with a conspicuous tubercle or “beak” (= hardened and<br />

persistent style base) at <strong>the</strong> apex (hence <strong>the</strong> name BEAK-RUSH).<br />

AA genus <strong>of</strong> over 250 species nearly cosmopolitan in distribution, with greatest diversity in <strong>the</strong><br />

New World tropics; <strong>the</strong>y are mostly <strong>plants</strong> <strong>of</strong> sunny places with wet, acidic soils (Kral 2002c).<br />

Temperate North America is rich in species, with ca. 60 in <strong>the</strong> se U.S. (Tucker 1987). According <strong>to</strong><br />

Yatskievych (1999), species <strong>of</strong> this genus “are generally considered poor forage for lives<strong>to</strong>ck. The<br />

tiny sharp teeth along <strong>the</strong> leaf margins are composed <strong>of</strong> silica, and <strong>the</strong>y make <strong>the</strong> <strong>plants</strong> relatively<br />

unfit for consumption.” However, many marshland species provide food for migra<strong>to</strong>ry waterfowl<br />

(Kral 2002c). In <strong>the</strong> key and descriptions presented here, <strong>the</strong> measurements given for achenes do<br />

not include <strong>the</strong> tubercles. (Greek: rhyncos, a snout, and spora, a seed, from <strong>the</strong> beaked achenes)<br />

REFERENCES: Gale 1944; Godfrey & Wooten 1979; Nixon & Ward 1982; Thomas 1984, 1992;<br />

Tucker 1987; Kral 1999, 2002c; Bridges & Orzell 2000.<br />

1. Scales <strong>of</strong> spikelets white; bracts below inflorescence with white base, <strong>the</strong> white zone conspicuous<br />

or inconspicuous; perianth bristles absent (section Dichromena, previously segregated as<br />

<strong>the</strong> genus Dichromena).<br />

2. Bracts exceeding spikelets 1–2(–3) in number, very narrow, mostly filiform; white zone on<br />

bract only at <strong>the</strong> very base, not longer than spikelets, inconspicuous; rhizomes absent; leaf<br />

blades 3–15 mm long, ca. 1(–2) mm broad basally, narrower (arcuate-filiform) distally ____________ R. nivea

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