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keys to the vascular plants of east texas - Botanical Research ...

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ERIONEURON/POACEAE<br />

945<br />

Shaw and Smeins (1983) concluded that it represents swollen or expanded internode tissue<br />

fused with <strong>the</strong> much-reduced lower glume. Thompson et al. (1990), however, found that it lacks<br />

<strong>vascular</strong> tissue, is formed entirely <strong>of</strong> parenchyma<strong>to</strong>us tissue, and that glume tissue is not involved.<br />

They concluded that it is “formed by a proliferation <strong>of</strong> <strong>the</strong> ground tissue at <strong>the</strong> spikelet<br />

base.” Davidse (1987a) suggested a possible function—<strong>the</strong> cup-shaped structure might contain<br />

lipids and function as an elaiosome <strong>to</strong> attract ants that would act as dispersal agents for <strong>the</strong><br />

seeds. (Greek: erion, wool, and chloa, grass, in reference <strong>to</strong> <strong>the</strong> densely hairy pedicels and spikelets<br />

<strong>of</strong> <strong>the</strong> type species—Shaw & Webster 1981) (subfamily Panicoideae, tribe Paniceae)<br />

REFERENCES: Shaw & Smeins 1979, 1981, 1983; Shaw & Webster 1987; Thompson et al. 1990;<br />

Crins 1991; Arriaga 2000; Shaw et al. 2003.<br />

1. Pedicels with long silky hairs ca. half as long as <strong>the</strong> spikelets or longer; inflorescence branches<br />

(racemes) not or only scarcely overlapping each o<strong>the</strong>r; leaf blades 0.5–4 mm wide; lemma <strong>of</strong><br />

perfect floret awnless or with a minute awn tip less than 0.4 mm long _________________________ E. sericea<br />

1. Pedicels glabrous or with only very short pubescence; inflorescence branches overlapping each<br />

o<strong>the</strong>r; leaf blades 2–10(–13) mm wide; lemma <strong>of</strong> perfect floret with an awn 0.4–1.5 mm long.<br />

2. Leaf blades glabrous or scabrous on lower surface; <strong>plants</strong> perennial (but may flower first year<br />

and appear annual); spikelets (4–)4.5–5.7 mm long, typically paired (but becoming solitary<br />

<strong>to</strong>ward tip <strong>of</strong> inflorescence branch); species known in East TX only from s portion <strong>of</strong> area<br />

_____________________________________________________________________________ E. punctata<br />

2. Leaf blades pubescent on lower surface; <strong>plants</strong> annual; spikelets 3.5–4.5(–5) mm long, solitary<br />

(occasionally paired at base <strong>of</strong> inflorescence branch); species widespread in East TX ________ E. contracta<br />

Eriochloa contracta Hitchc., (contracted), PRAIRIE CUP GRASS. Annual 15–75(–100) cm tall, <strong>of</strong>ten with<br />

many culms, forming bushy clumps; leaf sheaths and blades (at l<strong>east</strong> <strong>the</strong> lower ones) <strong>of</strong>ten ra<strong>the</strong>r<br />

densely s<strong>of</strong>t-pubescent; leaf blades 2–8 mm wide; lemma <strong>of</strong> perfect floret with awn 0.4–1.1 mm<br />

long; spikelets mostly solitary. Roadsides, ditches, disturbed areas; in East TX mainly Blackland<br />

Prairie, but widespread in TX; se Canada (Ont.) and sc and sw U.S., scattered elsewhere. May–Oct.<br />

Eriochloa punctata (L.) Desv. ex Hamil<strong>to</strong>n, (spotted), LOUISIANA CUP GRASS, EVERLASTING GRASS.<br />

Perennial, rhizoma<strong>to</strong>us, 30–150 cm tall; leaf sheaths and blades usually glabrous or blades scabrous;<br />

leaf blades (2–)4–10(–13) mm wide; lemma <strong>of</strong> perfect floret with awn 0.6–1.5 mm long;<br />

spikelets mostly paired. Ditches and o<strong>the</strong>r moist areas; Brazos and Harris (TAES) cos.; mainly<br />

Gulf Prairies and Marshes and South TX Plains; LA and TX. Mar–Dec. [Milium punctatum L.]<br />

This species is <strong>the</strong> l<strong>east</strong> common member <strong>of</strong> <strong>the</strong> genus in East TX. It is considered a significant<br />

weed in some areas (presumably where not native) by Watson and Dallwitz (1992).<br />

Eriochloa sericea (Scheele) Munro ex Vasey, (silky), TEXAS CUP GRASS, SILKY CUP GRASS. Perennial<br />

with short rhizomes but tufted, 30–130 cm tall, ra<strong>the</strong>r densely s<strong>of</strong>t-pubescent; spikelets 4–5<br />

mm long, solitary; lemma <strong>of</strong> perfect floret awnless or with a minute awn tip less than 0.4 mm<br />

long. Prairies, open thickets, roadsides, calcareous clays or clay loams; widespread in w part <strong>of</strong><br />

East TX from Leon (BRIT) and Grimes (Turner et al. 2003) cos. w; w portion <strong>of</strong> East TX w <strong>to</strong><br />

Rolling Plains and s <strong>to</strong> Edwards Plateau, South TX Plains, and Gulf Prairies and Marshes; CO,<br />

KS, NE, OK, and TX. May–Oct. This is a secondary member <strong>of</strong> <strong>the</strong> original prairie. It is reported<br />

<strong>to</strong> be good forage but does not withstand grazing; it is said <strong>to</strong> persist in overgrazed pastures only<br />

under <strong>the</strong> protection <strong>of</strong> shrubs (Hatch et al. 1999).<br />

ERIONEURON Nash WOOLLY GRASS, FLUFF GRASS<br />

AA New World C4 genus <strong>of</strong> 3 species (Valdés-Reyna & Hatch 1997; Valdés-Reyna 2003b);<br />

similar <strong>to</strong> and previously included in Tridens. Seedlings “appear <strong>to</strong> have a shaggy, white-villous<br />

‘indumentum,’ but this is composed <strong>of</strong> a myriad <strong>of</strong> small, water-soluble crystals” (Valdés-<br />

Reyna 2003b). The monotypic genus Dasyochloa <strong>of</strong> <strong>the</strong> sw U.S. and n Mexico has at times been

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