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keys to the vascular plants of east texas - Botanical Research ...

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420 AGAVACEAE/YUCCA<br />

Austin Chalk, Hill, McLennan (BRIT), Bell, Williamson, and Travis (Turner et al. 2003) cos.);<br />

mainly Cross Timbers and Prairies; endemic <strong>to</strong> TX (Kartesz 1999; Carr 2002b, 2002c). May–Jun.<br />

According <strong>to</strong> Clary (1997), DNA evidence suggests a close relationship <strong>of</strong> this species with Y.<br />

rupicola. Plants apparently intermediate between Y. pallida and Y. rupicola are found in several<br />

localities (McKelvey 1947), including <strong>the</strong> area n <strong>of</strong> Austin (E. Keith, pers. comm.). They usually<br />

have blue-green glaucous leaves (like Y. pallida), but <strong>the</strong> leaves are twisted and have wavy margins.<br />

E m/308<br />

Yucca rupicola Scheele, (growing on cliffs or ledges), TEXAS YUCCA, TWIST-LEAF YUCCA. Similar <strong>to</strong><br />

Y. pallida; colonies typically with 2–15 rosettes; leaves � 100 per rosette, 20–60 cm long, <strong>to</strong> 40<br />

mm wide, flaccid, twisted, <strong>the</strong> margins wavy; panicle glabrous <strong>to</strong> slightly pubescent; perianth<br />

segments whitish or greenish white; stigma lobes widely spreading. Limes<strong>to</strong>ne ledges, plains;<br />

Bexar, Comal, Hays, McLennan, Travis, and Williamson (Turner et al. 2003) cos. near w margin<br />

<strong>of</strong> <strong>the</strong> Blackland Prairie; mainly Edwards Plateau; endemic <strong>to</strong> TX (Kartesz 1999; Carr 2002b,<br />

2002c). Apr–Jun. E m/308<br />

Yucca <strong>to</strong>rreyi Shafer, (for John Torrey, 1796–1873, American botanist, physician, and collec<strong>to</strong>r <strong>of</strong><br />

many w North American <strong>plants</strong>), TORREY’S YUCCA, SPANISH-DAGGER. Plant large, <strong>to</strong> ca. 7 m tall;<br />

stems unbranched or rarely with 2–3 branches; dead leaves reflexed on trunk below leaf crown;<br />

leaves 30–110 cm long, light green, stiff, spear-like, entire, with marginal fibers, <strong>the</strong> apical portion<br />

<strong>of</strong> <strong>the</strong> leaves usually rolled inward so that <strong>the</strong> margins nearly <strong>to</strong>uch; panicle with 0.1–0.5 <strong>of</strong><br />

its <strong>to</strong>tal length extending beyond <strong>the</strong> leaves or rarely entirely within <strong>the</strong> leaves; perianth<br />

subglobose or campanulate, sometimes fully expanding, cream (can be tinged with purple);<br />

fruits 7–14 cm long, indehiscent, fleshy. Gravelly soils, grassy and chaparral mesas and slopes;<br />

Bexar (Turner et al. 2003) and Travis (Carr 2002a) cos. near <strong>the</strong> sw margin <strong>of</strong> East TX; also<br />

Brown and Burnet (HPC) cos. on sw margin <strong>of</strong> <strong>the</strong> Cross Timbers and Prairies; however, K. Clary<br />

(pers. comm.) questions whe<strong>the</strong>r <strong>the</strong> species is native as far e as Burnet and Travis cos.; mainly<br />

Edwards Plateau and Trans-Pecos. Late Mar–May. [Y. baccata Torr. var. macrocarpa Torr., Y.<br />

crassifolia Engelm., Y. macrocarpa (Torr.) Coville] This species is sometimes treated as a synonym<br />

<strong>of</strong> <strong>the</strong> related Y. treculeana (e.g., Powell 1988; Hess & Robbins 2002); according <strong>to</strong> Webber<br />

(1953) and Correll and Johns<strong>to</strong>n (1970), <strong>the</strong> two sometimes hybridize. Since <strong>the</strong>y can usually be<br />

readily distinguished in <strong>the</strong> field, we are following McKelvey (1938), Jones et al. (1997), and<br />

Kartesz (1999) in recognizing <strong>the</strong>m at <strong>the</strong> specific level.<br />

Yucca treculeana Carr., (for A.A.L. Trécul, 1818–1896, who <strong>to</strong>ok <strong>plants</strong> <strong>of</strong> this species <strong>to</strong> France in<br />

1850—Vines 1960), SPANISH-DAGGER, SPANISH-BAYONET, TRECUL’S YUCCA, DON QUIXOTE’S-LANCE,<br />

PITA, PALMA PITA, PALMA DE DÁTILES, PALMA LOCA, TEXAS-BAYONET. Plant large, <strong>to</strong> ca. 7 m tall;<br />

stems few-branched, with leaf crown at apex; dead leaves reflexed on trunk below leaf crown<br />

(trunks bare <strong>of</strong> dead leaves on old <strong>plants</strong>); leaves 50–100 cm long, green, entire (sometimes denticulate<br />

at base <strong>of</strong> leaf), without marginal fibers, <strong>the</strong> apical portions <strong>of</strong> <strong>the</strong> leaves not inrolled;<br />

panicle with ca. 0.5–0.75 <strong>of</strong> its <strong>to</strong>tal length extending beyond <strong>the</strong> leaves; perianth broadly globose<br />

or hemispherical, greenish cream <strong>to</strong> cream (can be lightly tinged with purple); fruits 5–11.5<br />

cm long, indehiscent, <strong>the</strong> flesh sweetish and succulent. Brushland; Bexar (TAES), Goliad<br />

(TAMU), Hays (TAMU), Travis (Tharp letter quoted in McKelvey 1938; R. O’Kennon, pers. obs.),<br />

and Comal (Turner et al. 2003) cos. near extreme sw margin <strong>of</strong> East TX, also Brazos Co. (K.<br />

Clary, pers. comm.); s TX n <strong>to</strong> s part <strong>of</strong> Cross Timbers and Prairies (Burnet Co.—Buckley in<br />

McKelvey 1938; R. O’Kennon, pers. obs.) and s part <strong>of</strong> East TX; in <strong>the</strong> U.S. known only from TX.<br />

Feb–Apr. [Y. treculeana var. succulenta McKelvey] Carr (2002a) considered Travis Co. collections<br />

previously thought <strong>to</strong> be <strong>of</strong> this species <strong>to</strong> be Y. <strong>to</strong>rreyi; however, K. Clary (pers. comm.) observes<br />

that Y. treculeana does come as far e as Travis Co. and also up from s TX <strong>to</strong> Brazos Co. The<br />

spines were used his<strong>to</strong>rically <strong>to</strong> jab a snake bite and induce bleeding in order <strong>to</strong> flush away <strong>the</strong><br />

poison (Vines 1960). Pioneers cooked and prepared <strong>the</strong> flowers like cabbage and also pickled<br />

<strong>the</strong>m (Schulz 1922; McKelvey 1938; Crosswhite 1980). According <strong>to</strong> Havard (1896), <strong>the</strong> fleshy,

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