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Final Comprehensive Conservation Plan - U.S. Fish and Wildlife ...

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Natural Processes<br />

(biotic <strong>and</strong> abiotic processes<br />

responsible for habitat conditions) Limiting Factors<br />

Collapse of lava tube roof <strong>and</strong> resulting Limiting factors<br />

vertical walled opening protects include a lack of native<br />

remnants of once common plants, now pollinators <strong>and</strong> several<br />

rare or endangered, from grazing by pest species (e.g.,<br />

ungulates.<br />

ungulates, rats, mice,<br />

slugs, mosquitoes,<br />

invasive plants, <strong>and</strong><br />

diseases).<br />

Population/Habitat Attributes<br />

(age class, structure, serial stage, species<br />

composition)<br />

Habitats (plant<br />

communities that represent<br />

existing BIDEH)<br />

The vertical walls created from the collapse of<br />

lava tube roofs naturally protect native plant<br />

communities. Skylights, unlike the lava tubes<br />

themselves, are exposed to the surface<br />

environment of rainfall, sunlight, <strong>and</strong><br />

temperature fluctuations.<br />

Lava tube skylights<br />

[Various above-ground<br />

habitats surround<br />

skylights, depending upon<br />

elevation]<br />

Hakalau Forest National <strong>Wildlife</strong> Refuge<br />

<strong>Comprehensive</strong> <strong>Conservation</strong> <strong>Plan</strong><br />

There are a finite<br />

number of lava tubes.<br />

Typically skylights<br />

form shortly after lava<br />

tube formation.<br />

These steep-sided depressions prevent<br />

degradation from ungulates, thus providing<br />

protected habitat for one fern species,<br />

Asplenium peruvianum var. insulare that is<br />

unique to skylight habitats, <strong>and</strong> also high<br />

populations of other rare <strong>and</strong> endangered<br />

plant species.<br />

Human disturbance<br />

such as trampling could<br />

destroy any sub-fossil<br />

or archeological<br />

resources.<br />

Appendix F: Biological Integrity, Diversity, <strong>and</strong> Environmental Health <strong>and</strong> Resources of Concern F-9<br />

Note: nomenclature for habitat names follows Jacobi et al. (1989).

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