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Maclean et al. - 2002 - Rice almanac source book for the most important e

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Flag leaf<br />

blade<br />

Axis<br />

Spikel<strong>et</strong><br />

(flor<strong>et</strong>)<br />

Secondary<br />

branch<br />

Primary branch<br />

Base<br />

Awn<br />

P<strong>al</strong>e<strong>al</strong> apiculus<br />

An<strong>the</strong>r<br />

Filament<br />

P<strong>al</strong>ea<br />

Lemma<br />

Stigma<br />

Style<br />

Ovary<br />

Rachilla<br />

Sterile lemmas<br />

Rudimentary glumes<br />

Pedicel<br />

Fig. 4. <strong>Rice</strong> panicle and spikel<strong>et</strong>s.<br />

and secondary branches; <strong>the</strong> spikel<strong>et</strong>s are<br />

positioned above <strong>the</strong>m (Fig. 4).<br />

Since rice has only one fully developed<br />

flor<strong>et</strong> (flower) per spikel<strong>et</strong>, <strong>the</strong>se terms are often<br />

used interchangeably. The flower is enclosed in<br />

<strong>the</strong> lemma and p<strong>al</strong>ea, which may be ei<strong>the</strong>r awned<br />

or awnless. The flower consists of <strong>the</strong> pistil and<br />

stamens, and <strong>the</strong> components of <strong>the</strong> pistil are <strong>the</strong><br />

stigmas, styles, and ovary.<br />

Growth<br />

The growth duration of <strong>the</strong> rice plant is 3–6<br />

months, depending on <strong>the</strong> vari<strong>et</strong>y and <strong>the</strong><br />

environment under which it is grown. During<br />

this time, rice compl<strong>et</strong>es two distinct growth<br />

phases: veg<strong>et</strong>ative and reproductive. The veg<strong>et</strong>ative<br />

phase is subdivided into germination, early<br />

seedling growth, and tillering; <strong>the</strong> reproductive<br />

phase is subdivided into <strong>the</strong> time be<strong>for</strong>e and after<br />

heading, that is, panicle exsertion. The time after<br />

heading is b<strong>et</strong>ter known as <strong>the</strong> ripening period<br />

(Fig. 5).<br />

Potenti<strong>al</strong> grain yield is primarily d<strong>et</strong>ermined<br />

be<strong>for</strong>e heading. Ultimate yield, which is based<br />

on <strong>the</strong> amount of starch that fills <strong>the</strong> spikel<strong>et</strong>s, is<br />

largely d<strong>et</strong>ermined after heading. Hence,<br />

agronomic<strong>al</strong>ly, it is convenient to regard <strong>the</strong> life<br />

history of rice in terms of three growth phases:<br />

veg<strong>et</strong>ative, reproductive, and ripening. A 120-d<br />

vari<strong>et</strong>y, when planted in a tropic<strong>al</strong> environment,<br />

spends about 60 d in <strong>the</strong> veg<strong>et</strong>ative phase, 30 d<br />

in <strong>the</strong> reproductive phase, and 30 d in <strong>the</strong><br />

ripening phase.<br />

Veg<strong>et</strong>ative phase<br />

The veg<strong>et</strong>ative phase is characterized by active<br />

tillering, a gradu<strong>al</strong> increase in plant height, and<br />

The rice plant and its ecology 13

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