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Maclean et al. - 2002 - Rice almanac source book for the most important e

Maclean et al. - 2002 - Rice almanac source book for the most important e

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engineering is being employed to introduce<br />

resistance to stem borers and sheath blight, two<br />

pests <strong>for</strong> which limited resistance has been found<br />

in rice germplasm. D<strong>et</strong>ails are given in <strong>the</strong><br />

section on “Biotechnology issues”.<br />

More sustainable integrated weed<br />

management technology is <strong>al</strong>so being developed<br />

by investigating and promoting rice cultivars<br />

with superior weed comp<strong>et</strong>itiveness, biologic<strong>al</strong><br />

control, rice <strong>al</strong>lelopathy, and a thorough<br />

understanding of <strong>the</strong> biology and ecology of<br />

weeds, and of <strong>the</strong> socioeconomics of weed<br />

management practices of farmers. Integrated<br />

weed management, promoting biologic<strong>al</strong> and<br />

cultur<strong>al</strong> control, and minimizing <strong>the</strong> use of<br />

herbicides are seen as keys to sustainable ricefarming<br />

systems.<br />

Friends<br />

As mentioned, <strong>most</strong> of <strong>the</strong> organisms in a rice<br />

field are not harmful. Control of insect pests, <strong>for</strong><br />

example, depends on <strong>the</strong> activities of a whole<br />

array of <strong>the</strong>ir natur<strong>al</strong> enemies, through a complex<br />

and rich food web of gener<strong>al</strong>ist and speci<strong>al</strong>ist<br />

predators and parasites that live above, below,<br />

and at <strong>the</strong> water surface as well as in flooded and<br />

aerated soil habitats (Fig. 8). In tropic<strong>al</strong> rice<br />

fields, levees or bunds <strong>al</strong>so <strong>for</strong>m refuges <strong>for</strong><br />

early season predators of rice pests, such as ants<br />

and spiders, as do bordering grasses and o<strong>the</strong>r<br />

nearby habitats.<br />

Predators are <strong>the</strong> <strong>most</strong> conspicuous group of<br />

rice “friends,” that is, <strong>the</strong> natur<strong>al</strong> enemies of rice<br />

pests. They are <strong>the</strong> <strong>most</strong> abundant <strong>for</strong>ms and are<br />

often confused with pests. Some, such as spiders,<br />

predatory crick<strong>et</strong>s, longhorned grasshoppers,<br />

damselflies, ants, wasps, and be<strong>et</strong>les, search <strong>the</strong><br />

plant <strong>for</strong> prey such as leafhoppers, planthoppers,<br />

moths, caterpillars, and larvae of stem borers.<br />

Some predators eat eggs of pests. O<strong>the</strong>rs, such as<br />

water bugs and be<strong>et</strong>les, live on <strong>the</strong> surface of and<br />

in <strong>the</strong> water in <strong>the</strong> rice field and attack pests that<br />

f<strong>al</strong>l from <strong>the</strong> plants.<br />

Canopy<br />

Neuston<br />

Plankton<br />

Epibenthos<br />

Benthos<br />

Fig. 8. Habitat zones and dominant taxa of an irrigated rice farmer’s rice field in C<strong>al</strong>auan, Laguna,<br />

Philippines. Canopy taxa are Stilbus sp. adults (1), Chironomidae adults (2), Cardiochiles philippinensis adults<br />

(3), and Egadroma sp. adults (4). Neustonic (water surface) taxa are Microvelia atrolineata adults (5),<br />

Ephydra sp. immatures (6), Entomobryidae adults (7), and M. atrolineata immatures (8). Planktonic and<br />

benthic taxa are H<strong>et</strong>erocypris luzonensis (9), Eucyclops serrulatus adults (10), Chironomidae immatures (11),<br />

and Hydrophilidae adults (12).<br />

28 <strong>Rice</strong> <strong>al</strong>manac

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