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Cambridge International A Level Biology Revision Guide

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Chapter 12: Energy and respiration<br />

CO 2 + H 2 O<br />

respiration<br />

chemical potential<br />

energy of ATP<br />

Key<br />

transfer of materials<br />

transfer of energy<br />

Figure 12.2 Transfer of materials and energy in an ecosystem.<br />

Work<br />

Work in a living organism includes:<br />

■■<br />

■■<br />

■■<br />

■■<br />

light energy<br />

autotrophs<br />

e.g. green plants<br />

heterotrophs<br />

e.g. animals, fungi<br />

and most bacteria<br />

chemical potential<br />

energy of carbohydrates<br />

and other organic<br />

molecules<br />

thermal energy<br />

work<br />

the synthesis of complex substances from simpler<br />

ones (anabolic reactions), such as the synthesis of<br />

polysaccharides from monosaccharides, lipids from<br />

glycerol and fatty acids, polypeptides from amino<br />

acids, and nucleic acids from nucleotides<br />

the active transport of substances against a diffusion<br />

gradient, such as the activity of the sodium–potassium<br />

pump (Figure 4.18, page 87)<br />

mechanical work such as muscle contraction<br />

(page 344) and other cellular movements; for example,<br />

the movement of cilia and flagella (page 189), amoeboid<br />

movement and the movement of vesicles through<br />

cytoplasm<br />

in a few organisms, bioluminescence and<br />

electrical discharge.<br />

Mammals and birds use thermal energy (heat) that is<br />

released from metabolic reactions to maintain a constant<br />

body temperature. Most animals are ectotherms. The<br />

thermal energy that warms them comes from outside their<br />

bodies. Mammals and birds are endotherms, releasing<br />

enough thermal energy within their bodies to maintain<br />

them above the temperature of their surroundings<br />

when necessary. They also maintain a constant body<br />

temperature through negative feedback loops (page 301).<br />

For a living organism to do work, energy-requiring<br />

reactions must be linked to those that yield energy. In<br />

the complete oxidation of glucose (C 6<br />

H 12<br />

O 6<br />

) in aerobic<br />

conditions, a large quantity of energy is made available:<br />

C 6<br />

H 12<br />

O 6<br />

+ 6O 2<br />

→ 6CO 2<br />

+ 6H 2<br />

O + 2870 k J<br />

Reactions such as this take place in a series of small<br />

steps, each releasing a small quantity of the total available<br />

energy. Multi-step reactions allow precise control via<br />

feedback mechanisms (Chapter 3). Moreover, the cell could<br />

not usefully harness the total available energy if all of it<br />

were made available at one instant.<br />

Although the complete oxidation of glucose to carbon<br />

dioxide and water has a very high energy yield, the<br />

reaction does not happen easily. Glucose is actually quite<br />

stable, because of the activation energy that has to be<br />

added before any reaction takes place (Figure 12.3). In<br />

living organisms, the activation energy is overcome by<br />

lowering it using enzymes (page 56), and also by raising<br />

the energy level of the glucose by phosphorylation<br />

(page 272).<br />

Theoretically, the energy released from each step of<br />

respiration could be harnessed directly to some form of<br />

work in the cell. However, a much more flexible system<br />

actually occurs in which energy-yielding reactions in<br />

all organisms are used to make an intermediary<br />

molecule, ATP.<br />

Increase in energy<br />

C 6 H 12 O 6 + O 2<br />

substrates<br />

Progress of reaction<br />

Figure 12.3 Oxidation of glucose.<br />

CO 2 + H 2 O<br />

products<br />

activation<br />

energy<br />

available<br />

energy<br />

269

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