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Cambridge International A Level Biology Revision Guide

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Chapter 14: Homeostasis<br />

Stomata show daily rhythms of opening and closing.<br />

Even when kept in constant light or constant dark, these<br />

rhythms persist (Figure 14.30). Opening during the day<br />

maintains the inward diffusion of carbon dioxide and<br />

the outward diffusion of oxygen. However, it also allows<br />

the outward diffusion of water vapour in transpiration<br />

(pages 134 –136). The closure of stomata at night when<br />

photosynthesis cannot occur reduces rates of transpiration<br />

and conserves water.<br />

Stomata respond to changes in environmental<br />

conditions. They open in response to:<br />

■■<br />

■■<br />

increasing light intensity<br />

low carbon dioxide concentrations in the air spaces<br />

within the leaf.<br />

When stomata are open, leaves gain carbon dioxide<br />

for photosynthesis, but tend to lose much water in<br />

transpiration.<br />

a<br />

6<br />

4<br />

2<br />

b<br />

0<br />

c<br />

Stomatal opening / arbitrary units<br />

6<br />

4<br />

2<br />

0<br />

321<br />

6<br />

4<br />

2<br />

0<br />

24 12 24 12 24 12 24 12 24 12<br />

Time of day (12 = midday, 24 = midnight,<br />

= darkness)<br />

Figure 14.30 a The opening of stomata was measured in leaves of Tradescantia over several days to reveal a<br />

daily rhythm of opening and closing. This rhythm persisted even when the plants were kept in constant light b;<br />

and in long periods of constant dark c.

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