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Cambridge International A Level Biology Revision Guide

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Chapter 18: Biodiversity, classification and conservation<br />

100 smaller squares. You then decide approximately what<br />

percentage of the area inside the quadrat is occupied by<br />

each species. These percentages may not add up to 100%.<br />

For example, there might be bare ground in the quadrat,<br />

so the numbers will come to less than 100%. Or there<br />

may be plants overlying one another, in which case the<br />

numbers may add up to more than 100%. An alternative<br />

to estimating percentage cover of each species is to use<br />

an abundance scale, such as the Braun–Blanquet scale for<br />

number and plant cover (Table 18.1).<br />

QUESTIONS<br />

18.4 A survey gave the following results for a species<br />

of the red sea anemone, Isactinia tenebrosa<br />

(Figure 18.10), on a rocky shore in New Zealand,<br />

using a quadrat with an area of 0.25 m 2 .<br />

Quadrat 1 2 3 4 5 6 7 8 9 10<br />

Number<br />

of red sea<br />

anemones<br />

0 3 0 1 0 0 5 2 0 1<br />

a Calculate i the species frequency, and ii the<br />

species density of I. tenebrosa from the results of<br />

this survey.<br />

b Suggest when it might be more appropriate to<br />

use species frequency rather than species density<br />

to record the abundance of a species.<br />

18.5 A survey was made of Benghal dayflower,<br />

Commelina benghalensis, growing on a lawn and in<br />

a field of young soybean plants. Ten 1.0 m 2 quadrats<br />

were placed randomly in each area, and the number<br />

of dayflower plants in each quadrat was counted.<br />

The results are shown in the table.<br />

Quadrat 1 2 3 4 5 6 7 8 9 10<br />

Number of<br />

dayflowers<br />

on lawn<br />

0 0 4 3 0 1 2 4 0 3<br />

429<br />

Figure 18.9 Estimating percentage cover. This 1 m 2 quadrat<br />

is divided into 100 small squares to make it easier to make the<br />

estimation for each species.<br />

Number of<br />

dayflowers<br />

in field<br />

0 0 0 2 5 0 0 1 0 0<br />

Braun–Blanquet Cover Scale<br />

Description<br />

Value<br />

Very few plants, cover is less than 1% +<br />

Many plants, but cover is 1–5% 1<br />

Very many plants or cover is 6–25% 2<br />

Any number of plants; cover is 26–50% 3<br />

Any number of plants; cover is 51–75% 4<br />

Cover is greater than 75% 5<br />

a Calculate i the species frequency, and ii the<br />

species density of dayflower plants in each of the<br />

two areas.<br />

b Explain why it is important to use randomly<br />

placed quadrats.<br />

c Suggest two disadvantages with calculating<br />

percentage cover or using an abundance scale,<br />

such as the Braun–Blanquet scale.<br />

Table 18.1 The Braun–Blanquet scale for recording<br />

vegetation within quadrats.<br />

Figure 18.10 A red sea anemone that lives between the tides<br />

on rocky shores.

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