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Principles of Plant Genetics and Breeding

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Taxonomy<br />

Kingdom <strong>Plant</strong>ae<br />

Subkingdom Tracheobionta<br />

Superdivision Spermatophyta<br />

Division Magnoliophyta<br />

Class Lilliopsida<br />

Subclass Commelinidae<br />

Order Cyperales<br />

Family Poaceae<br />

Genus Oryza L.<br />

Species Oryza sativa L.<br />

Economic importance<br />

Rice accounts for about 20% <strong>of</strong> the world’s total grain<br />

production, second only to wheat. It is the primary<br />

staple for more than 50% <strong>of</strong> the world’s population.<br />

An estimated 90% <strong>of</strong> production <strong>and</strong> consumption <strong>of</strong><br />

rice occurs in Asia where the per capita consumption<br />

is about 45 kg, compared to a world average <strong>of</strong> about<br />

27 kg <strong>and</strong> a US average <strong>of</strong> about 9 kg. World consumption<br />

in 1993/94 was 358.5 million metric tons <strong>and</strong> has<br />

increased steadily to an estimated 408.8 million tons in<br />

2002/03. Similarly, the area devoted to rice increased<br />

during the same period from 145.2 to 155.1 ha in<br />

1990/2000, with a slight drop thereafter. Production<br />

trends mirror the area cultivated. Rice production in the<br />

USA is dominated by six states – Arkansas, California,<br />

Louisiana, Texas, Mississippi, <strong>and</strong> Missouri – together<br />

accounting for about 99% <strong>of</strong> the total US production.<br />

Other minor producers in the USA include Florida,<br />

Tennessee, Illinois, South Carolina, <strong>and</strong> Kentucky.<br />

Arkansas leads the production in the US, with about<br />

100 cwt <strong>of</strong> rice produced in 2001 on about 1.61 million<br />

acres (see Appendix 2 for conversion rates for units).<br />

The world’s major producers <strong>of</strong> rice are China, India,<br />

Indonesia, <strong>and</strong> Bangladesh, which together account for<br />

29<br />

<strong>Breeding</strong> rice<br />

more than 70% <strong>of</strong> the world’s total production. China,<br />

the world’s leading producer regularly accounts for<br />

about 36% <strong>of</strong> the world’s total production. The expansion<br />

in rice production is attributed largely to the impact<br />

<strong>of</strong> the Green Revolution, which was implemented in the<br />

1960s <strong>and</strong> 1970s in Asia <strong>and</strong> other parts <strong>of</strong> the tropical<br />

world. Other major producers in Asia include Vietnam,<br />

Thail<strong>and</strong>, Japan, <strong>and</strong> Burma, together accounting for<br />

about 12% <strong>of</strong> the total world production. Outside <strong>of</strong><br />

Asia, rice is produced in substantial amounts in countries<br />

including the USA, Italy, Spain, Egypt, Australia,<br />

<strong>and</strong> countries in the West African region.<br />

Thail<strong>and</strong> leads the world in rice exports, accounting<br />

for about 25% <strong>of</strong> the total world exports (about 4 million<br />

metric tons). The USA is the second largest<br />

exporter <strong>of</strong> rice, exporting about 2.7 million metric<br />

tons. Latin America is the largest market for US rice<br />

export. Other major exporters <strong>of</strong> rice include Vietnam,<br />

Pakistan, China, Australia, Burma, Italy, India, <strong>and</strong><br />

Uruguay.<br />

Origin <strong>and</strong> history<br />

The origin <strong>of</strong> rice is not exactly known. Wild species <strong>of</strong><br />

rice occur across South <strong>and</strong> East Asia, including India<br />

<strong>and</strong> south China. Rice was domesticated in the 5th<br />

millennium bc. The cultivated species <strong>of</strong> rice Oryza<br />

sativa is believed to have derived from annual progenitors<br />

found in a wide area extending from the Gangetic<br />

Plains, through Burma, northern Thail<strong>and</strong>, northern<br />

Vietnam, <strong>and</strong> southern China. Rice trade occurred<br />

between Egypt, India, <strong>and</strong> China. The Moors are credited<br />

with bringing rice to Spain, from where it was<br />

introduced to Italy (in the 15th century) <strong>and</strong> subsequently<br />

to Central America. The African cultigen<br />

(O. glaberrima) originated in the Niger River delta <strong>of</strong><br />

Mali in West Africa.

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