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Principles of Plant Genetics and Breeding

Principles of Plant Genetics and Breeding

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Purpose <strong>and</strong> expected outcomes<br />

<strong>Plant</strong> breeders manipulate the genotype <strong>of</strong> plants to create heritable modifications in plant structure <strong>and</strong> function.<br />

Sometimes plants are genetically modified to change their shape, height, or size, to facilitate a production operation<br />

or system. For example, dwarf plants are more environmentally responsive <strong>and</strong> tend to resist lodging. Changes in<br />

stature <strong>and</strong> size impact their metabolic activities. To undertake such modifications, it is important for breeders to<br />

underst<strong>and</strong> the fundamental plant structure <strong>and</strong> organization at the molecular, cellular, <strong>and</strong> whole-plant levels.<br />

Manipulating plants by some biotechnological procedures occurs at the cellular level, hence the need to underst<strong>and</strong><br />

DNA <strong>and</strong> cellular structure <strong>and</strong> function. Because the goal <strong>of</strong> plant breeding is to create modifications that are<br />

permanent <strong>and</strong> heritable, it is important to also underst<strong>and</strong> the genetic architecture <strong>of</strong> plants <strong>and</strong> how genes condition<br />

plant traits. This chapter is designed to present an overview <strong>of</strong> concepts pertaining to the cellular organization<br />

<strong>and</strong> genetic structure <strong>of</strong> plants. After studying this chapter, the student should be able to:<br />

1 Briefly describe plant cell structure <strong>and</strong> organization.<br />

2 Briefly discuss nuclear division processes.<br />

3 Discuss Mendelian concepts.<br />

4 Discuss DNA structure <strong>and</strong> function.<br />

5 Distinguish between phenotype <strong>and</strong> genotype.<br />

6 Discuss the role <strong>of</strong> genetic linkage in plant breeding.<br />

7 More importantly, the student should be able to discuss the role <strong>of</strong> these plant structures <strong>and</strong> processes in plant<br />

breeding.<br />

Units <strong>of</strong> organization <strong>of</strong> living things<br />

The cell is the fundamental unit <strong>of</strong> organization <strong>of</strong> living<br />

things. Some organisms consist entirely <strong>of</strong> one cell<br />

(called unicellular) whereas others consist <strong>of</strong> numerous<br />

cells working together (called multicellular). Except<br />

for a bacterium, which lacks cellular compartmentalization<br />

into discrete functional units called organelles, <strong>and</strong><br />

is called a prokaryote, all other cells have a membrane-<br />

3<br />

<strong>Plant</strong> cellular<br />

organization <strong>and</strong> genetic<br />

structure: an overview<br />

bound nucleus <strong>and</strong> several other membrane-enclosed<br />

organelles <strong>and</strong> are called eukaryotes.<br />

The cell can be a unit for selection in breeding if, for<br />

example, molecular tools are used. The technology <strong>of</strong><br />

genetic engineering targets single cells for manipulation.<br />

After successfully transferring foreign genes into<br />

the cell, it is isolated <strong>and</strong> nurtured into a full plant. On<br />

the other h<strong>and</strong>, when conventional tools are used, the<br />

whole plant is the unit <strong>of</strong> selection. It should be pointed

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