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Principles of Plant Genetics and Breeding

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Purpose <strong>and</strong> expected outcomes<br />

Climate is the variation in meteorological factors over a large area formed over many years. It determines plant<br />

adaptation to a growing region. Weather on the other h<strong>and</strong> is the environmental condition described by the shortterm<br />

variations in meteorological factors within a local area. It determines crop development <strong>and</strong> productivity.<br />

Weather affects the choice <strong>of</strong> specific crop production activities (i.e., how the crop is produced in the field). <strong>Plant</strong><br />

breeders usually develop breeding programs to produce cultivars for specific production regions. Crop production<br />

is subject to the vagaries <strong>of</strong> the weather. Unpredictable weather can drastically reduce crop yield. Crop varieties<br />

used in regions that are prone to adverse weather during production need to have the capacity to resist or tolerate<br />

environmental stresses to an extent that they produce acceptable economic yield. After completing this chapter, the<br />

student should be able to:<br />

1 Describe the types <strong>of</strong> environmental stresses that reduce plant performance in crop production.<br />

2 Discuss breeding for drought resistance.<br />

3 Discuss breeding for cold tolerance.<br />

4 Discuss breeding for salt tolerance.<br />

5 Discuss breeding for resistance to heat stress.<br />

6 Discuss breeding for tolerance to aluminum toxicity.<br />

7 Discuss breeding for tolerance to oxidative stress.<br />

8 Discuss breeding for resistance to water-logging.<br />

Importance <strong>of</strong> breeding for resistance to<br />

abiotic stresses<br />

Only about 30% <strong>of</strong> the earth is l<strong>and</strong>. Of this, about 50%<br />

is not suitable for economic crop production, mainly<br />

because <strong>of</strong> constraints <strong>of</strong> temperature, moisture, <strong>and</strong><br />

topography. Of the remaining portion <strong>of</strong> arable l<strong>and</strong>,<br />

optimum production is further limited by a variety <strong>of</strong><br />

environmental stresses, requiring mineral <strong>and</strong> moisture<br />

supplementation for economic crop production. As the<br />

world population increases more food will have to be<br />

produced by increasing the productivity <strong>of</strong> existing<br />

farml<strong>and</strong>s as well as bringing new l<strong>and</strong>s into production.<br />

21<br />

<strong>Breeding</strong> for resistance to<br />

abiotic stresses<br />

This means, marginal l<strong>and</strong>s will have to be considered.<br />

<strong>Plant</strong>s breeders will have to develop cultivars that are<br />

adapted to specific environmental stresses. It is estimated<br />

that abiotic environmental stresses are responsible for<br />

about 70% <strong>of</strong> yield reduction <strong>of</strong> crops in production.<br />

In modern agricultural production, producers (especially<br />

those in developed countries) are able to alleviate<br />

moisture stress by providing supplemental irrigation.<br />

Also, fertilizers are used to alleviate nutritional stress,<br />

while soil reaction (pH) can be modified by applying<br />

soil amendments such as lime or sulfur. Excessive soil<br />

moisture can be removed by installing drainage systems.<br />

However, in poor regions <strong>of</strong> the world where such

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