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Principles of Plant Genetics and Breeding

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104 CHAPTER 6<br />

<strong>and</strong> Agricultural Organization (FAO) <strong>of</strong> the United<br />

Nations (UN) is credited with the initial efforts to promote<br />

genetic conservation, <strong>and</strong> assistance in establishing<br />

the International Board <strong>of</strong> <strong>Plant</strong> Genetic Resources<br />

(IBPGR) based in the FAO in Rome, Italy. Founded<br />

in 1974, the IBPGR is funded by donor countries,<br />

development banks, <strong>and</strong> foundations. It is a center in<br />

the Consultative Group <strong>of</strong> International Agriculture<br />

Research (CGIAR). The primary role <strong>of</strong> this board is to<br />

collect, preserve, evaluate, <strong>and</strong> assist with the exchange<br />

<strong>of</strong> plant genetic material for specific crops all over the<br />

world.<br />

A major sponsor <strong>of</strong> these genetic conservation activities<br />

is the International Agricultural Research Centers<br />

(IARCs) strategically located throughout the tropics<br />

(see Chapter 25). Gene banks at these centers focus on<br />

starchy crops that feed the world (wheat, corn, rice,<br />

potato, sorghum). These crops are <strong>of</strong>ten grown with<br />

high-tech cultivars that have narrow genetic bases as a<br />

result <strong>of</strong> crop improvement.<br />

There are other regional- <strong>and</strong> country-based plant<br />

germplasm conservation programs. The EUCARPIA<br />

(European Association for <strong>Plant</strong> <strong>Breeding</strong> Research),<br />

started in 1960, serves Europe <strong>and</strong> the Mediterranean<br />

region. Similarly, the Vegetable Gene Bank at the National<br />

Vegetable Research Station in the UK was established in<br />

1981 to conserve vegetable genetic resources.<br />

Palmer<br />

Hilo<br />

Western<br />

North Central<br />

North Eastern<br />

Southern<br />

Corvallis<br />

Davis<br />

Parlier<br />

Riverside<br />

Pullman<br />

Aberdeen<br />

Ft. Collins<br />

Figure 6.2 The four regional germplasm jurisdictions defined by the USDA.<br />

An example <strong>of</strong> a national germplasm<br />

conservation system<br />

The US plant genetic conservation efforts are coordinated<br />

by the National <strong>Plant</strong> Germplasm System<br />

(NPGS). Over 400,000 accessions exist in the inventory<br />

<strong>of</strong> the NPGS in the form <strong>of</strong> seed <strong>and</strong> vegetative material.<br />

In August, 2004, the composition <strong>of</strong> the holdings was<br />

205 families, 1,644 genera, <strong>and</strong> 10,205 species, for a<br />

total <strong>of</strong> 460,799 accessions. Most <strong>of</strong> these materials are<br />

predominantly l<strong>and</strong>races <strong>and</strong> unimproved germplasm<br />

from overseas sources. The accessions in the NPGS are<br />

estimated to increase at a rate <strong>of</strong> 7,000–15,000 new<br />

entries per year. The system has certain component<br />

units with specific functions as follows.<br />

<strong>Plant</strong> introduction<br />

Located in Beltsville, Maryl<strong>and</strong>, the <strong>Plant</strong> Introduction<br />

Office is part <strong>of</strong> the <strong>Plant</strong> <strong>Genetics</strong> <strong>and</strong> Germplasm<br />

Institute <strong>of</strong> the USDA-ARS. Each entry is given a plant<br />

introduction (PI) number, but this unit does not maintain<br />

any plant material collection. The responsibilities<br />

to maintain, evaluate, <strong>and</strong> release plant materials are<br />

assigned to four regional <strong>Plant</strong> Introduction Stations<br />

(Western, North Central, North Eastern, <strong>and</strong> Southern)<br />

(Figure 6.2). The <strong>Plant</strong> Quarantine Facility <strong>of</strong> USDA<br />

Sturgeon Bay<br />

Ames<br />

Urbana<br />

College Station<br />

Columbus<br />

Griffin<br />

Geneva<br />

Beltsville<br />

Washington, DC<br />

Oxford<br />

Miami<br />

Mayaguez, PR

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