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Principles of Plant Genetics and Breeding

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PERFORMANCE EVALUATION FOR CROP CULTIVAR RELEASE 427<br />

Exp. Number 84HT01 2004 WHEAT HEADROWS<br />

Seed Source Cross Gen Parent Lines Code Plot No<br />

Check variety = Redcoat ****** 101<br />

83GH01 4 T830001 F1 FRANKENMUTH/AUGUSTA PMHFLR 102<br />

83GH01 4 103<br />

83GH01 6 T830002 F1 CALDWELL/AUGUSTA//HOUSER LRPM 104<br />

83GH01 6 105<br />

Figure 2 Output <strong>of</strong> a plant breeding book generated with BRSERIES in MSTAT.<br />

The use <strong>of</strong> MSTAT strengthens plant breeding programs in several ways: (i) by quickly <strong>and</strong> accurately designing yield trials <strong>and</strong><br />

other experiments; (ii) by enabling breeders to maintain better records; (iii) by increasing the size <strong>of</strong> the breeding programs<br />

because <strong>of</strong> the ease <strong>of</strong> information h<strong>and</strong>ling; (iv) by creating books with information collected over several seasons; <strong>and</strong> (v) by<br />

analyzing experimental data over multiple locations <strong>and</strong> years. MSTAT helps to improve the efficiency, timeliness, <strong>and</strong><br />

confidence <strong>of</strong> conclusions that are all very important concerns for plant breeders.<br />

interaction (AMMI). These procedures were discussed<br />

in Chapter 7.<br />

Adaptation<br />

According to P. M. A. Tigerstedt, in the context <strong>of</strong><br />

evolutionary biology, adaptation is a process, adaptiveness<br />

is the level <strong>of</strong> adaptation <strong>of</strong> a plant material to an<br />

environment, while adaptability is the ability to show<br />

adaptiveness in a wide range <strong>of</strong> environments. However,<br />

according to Cooper <strong>and</strong> Byth, in a plant breeding<br />

context, adaptation <strong>and</strong> adaptiveness relate to a condition<br />

rather than a process. They indicate the ability <strong>of</strong> a<br />

plant material to be high yielding with respect to a<br />

specific environment, one to which it is adapted. One<br />

<strong>of</strong> the purposes <strong>of</strong> a G × E analysis is to help breeders<br />

decide whether to breed for narrow or wide adaptation<br />

or adaptability. <strong>Breeding</strong> for narrow adaptability, the<br />

breeder’s goal is to release a cultivar for a specific part<br />

<strong>of</strong> the target region (with a unique set <strong>of</strong> conditions),<br />

whereas in breeding for wide adaptability, the breeder<br />

focuses on releasing a cultivar with high performance<br />

across all environments.<br />

As previously stated, only a G × location interaction<br />

is useful for determining genotype adaptation patterns<br />

in an ANOVA analysis. Further, only repeatable G ×<br />

location interactions are <strong>of</strong> practical importance. However,<br />

if the G × location variance is small in magnitude<br />

compared to other sources <strong>of</strong> variance, even though<br />

significant, it minimizes the specific advantage for<br />

breeding for narrow adaptation.<br />

Field plot technique in plant breeding<br />

The subject <strong>of</strong> field plot technique deals with decisionmaking<br />

processes <strong>and</strong> other activities conducted by the<br />

plant breeder to fairly <strong>and</strong> effectively evaluate genotypes<br />

from a breeding program, to select cultivars for release.<br />

Field trials are designed to evaluate genotypes in a<br />

natural environment that is similar to that which farmers<br />

operate in. To evaluate the genetic potential <strong>of</strong> genotypes,<br />

they must be grown in an identical environment<br />

so that any observed differences can be attributed to<br />

the genotype not chance variations. Unfortunately,<br />

outside a controlled environment condition such as a<br />

greenhouse, it is impossible to find a truly homogeneous<br />

environment in the field. One <strong>of</strong> the goals <strong>of</strong> the field<br />

plot technique is to evaluate genotypes error free (or,<br />

in practice, with a minimum error). It is important,<br />

therefore, to identify the source <strong>of</strong> error in a field study.<br />

In order to collect data in the proper manner for analysis<br />

<strong>and</strong> decision-making, the breeder should follow a<br />

set <strong>of</strong> statistically based rules for laying out an experiment<br />

in the field, called the experimental design. Such<br />

a design not only allows the breeder to estimate r<strong>and</strong>om<br />

error in the field but the choice <strong>of</strong> a good design can<br />

minimize this error.<br />

Sources <strong>of</strong> experimental error<br />

In an experiment, a researcher may deliberately impose<br />

conditions or factors that would generate variance in<br />

outcomes; such a factor is called a treatment. The unit<br />

to which a treatment is applied is the experimental

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