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Principles of Plant Genetics and Breeding

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Purpose <strong>and</strong> expected outcomes<br />

As previously indicated, plant breeders depend on variability for crop improvement. <strong>Plant</strong> genetic resources (plant<br />

germplasm) used in plant breeding are natural resources that are susceptible to erosion from use <strong>and</strong> abuse. It is<br />

important that they be collected, properly used, managed, <strong>and</strong> conserved to avoid irreparable loss <strong>of</strong> precious genetic<br />

material. After completing this chapter, the student should be able to:<br />

1 Discuss the importance <strong>of</strong> germplasm to plant breeding.<br />

2 Define the types <strong>of</strong> germplasm.<br />

3 Discuss the sources <strong>of</strong> germplasm for plant breeding.<br />

4 Discuss the mechanisms for conservation <strong>of</strong> germplasm.<br />

5 Discuss the international role in germplasm conservation.<br />

Importance <strong>of</strong> germplasm to plant breeding<br />

Germplasm is the lifeblood <strong>of</strong> plant breeding without<br />

which breeding is impossible to conduct. It is the genetic<br />

material that can be used to perpetuate a species<br />

or population. It not only has reproductive value, but<br />

through genetic manipulation (plant breeding), germplasm<br />

can be improved for better performance <strong>of</strong> the<br />

crop. Germplasm provides the materials (parents) used<br />

to initiate a breeding program. Sometimes, all plant<br />

breeders do is to evaluate plant germplasm <strong>and</strong> make a<br />

selection from existing biological variation. Promising<br />

genotypes that are adapted to the production region are<br />

then released to producers. Other times, as discussed in<br />

Chapter 5, breeders generate new variability by using a<br />

variety <strong>of</strong> methods such as crossing parents, mutagenesis<br />

(inducing mutations), <strong>and</strong>, more recently, gene transfer.<br />

This base population is then subjected to appropriate<br />

selection methods, leading to the identification <strong>and</strong> further<br />

evaluation <strong>of</strong> promising genotypes for realease as<br />

cultivars. When breeders need to improve plants, they<br />

6<br />

<strong>Plant</strong> genetic resources<br />

for plant breeding<br />

have to find a source <strong>of</strong> germplasm that would supply<br />

the genes needed to undertake the breeding project.<br />

To facilitate the use <strong>of</strong> germplasm, certain entities<br />

(germplasm banks) are charged with the responsibility<br />

<strong>of</strong> assembling, cataloguing, storing, <strong>and</strong> managing large<br />

numbers <strong>of</strong> germplasm. This strategy allows scientists<br />

ready <strong>and</strong> quick access to germplasm when they need it.<br />

Centers <strong>of</strong> diversity in plant breeding<br />

The subject <strong>of</strong> centers <strong>of</strong> diversity was first discussed<br />

in Chapter 2. Whereas the existence <strong>of</strong> centers <strong>of</strong><br />

crop origin or domestication is not incontrovertible, the<br />

existence <strong>of</strong> natural reservoirs <strong>of</strong> plant genetic variability<br />

has been observed to occur in certain regions <strong>of</strong> the<br />

world. These centers are important to plant breeders<br />

because they represent pools <strong>of</strong> diversity, especially wild<br />

relatives <strong>of</strong> modern cultivars.<br />

<strong>Plant</strong> breeding may be a victim <strong>of</strong> its own success.<br />

The consequence <strong>of</strong> selection by plant breeders in their

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