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PE EIE[R-Rg RESEARCH ON - HJ Andrews Experimental Forest

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have been developed . Prior to their development<br />

the overall ecosystem model will be for<br />

the most part a translation of simple graphica l<br />

and verbal models into mathematical language<br />

with many aspects of the system interpolate d<br />

on the basis of empirical submodels or eve n<br />

crude estimates .<br />

Ultimately both total systems models an d<br />

submodels will require testing. This testing<br />

will indicate the further research needed to<br />

answer new questions or refine parts of existing<br />

models . The modeling team plays a role in<br />

assisting in the design of such new research. I n<br />

this way the model building plays an essentia l<br />

feedback role in the program .<br />

In addition, both subsystem models an d<br />

larger ecosystem models require estimates of<br />

parameters and available data often appear s<br />

inadequate. In these circumstances, it is neces -<br />

sary to test the sensitivity of the model to th e<br />

errors of estimation. Does it make a great deal<br />

of difference if the estimates are crude? If<br />

not, the model may proceed . If it does make a<br />

significant difference, additional observation s<br />

are required and further research in such area s<br />

is necessary. Obviously, such sensitivity tests<br />

cannot be made until models are fairly wel l<br />

developed ; some models will be at this stage<br />

this year.<br />

Specific Goals<br />

The goal of our Coniferous Biome research<br />

is development of a basic understanding of<br />

coniferous forest ecosystems, including both<br />

terrestrial and aquatic components, so tha t<br />

ecological constraints on and opportunitie s<br />

for increased production of fiber, food, water ,<br />

and wildlife can be recognized . The overall<br />

strategy includes identification of the majo r<br />

components and processes, both physical an d<br />

organic, within the ecosystem, and definition<br />

of their interrelationships . The definition of<br />

interrelationships will be accomplished<br />

through a systems analysis and modeling pro -<br />

gram . As an overall guide for Biome research ,<br />

we recognize the following general objectives :<br />

1. To determine the major factors, both components<br />

and processes, that control the<br />

productivity and distribution of organisms<br />

in coniferous forest ecosystems, includin g<br />

(a) an analysis of the structure and distribution<br />

of the principal resources, (b) definition<br />

of the functional relationships between<br />

biotic, decomposer, consumer, and<br />

producer components of the systems, an d<br />

(c) analysis of the forms and degrees of<br />

stability in these systems .<br />

2. To examine the linkage of terrestrial an d<br />

aquatic components in coniferous fores t<br />

ecosystems, including (a) water, energy ,<br />

and transport of chemicals (including pesticides),<br />

(b) direct transport of terrestrial<br />

products into the aquatic system, e .g. ,<br />

through litter fall and surface erosion, an d<br />

(c) return of organic and inorganic materia l<br />

from the aquatic to the terrestrial environment<br />

through movements of fish, birds ,<br />

and insects .<br />

3. To determine how various manipulation s<br />

influence the structure and function of<br />

coniferous forest ecosystems using bot h<br />

unit watersheds and plot studies . Special<br />

attention is directed to the influences of<br />

manipulations on (a) stability and productivity<br />

of these systems and (b) the linkage s<br />

between terrestrial and aquatic components<br />

of the systems.<br />

4. To understand population dynamics of<br />

those major components of each trophic<br />

level which appear to influence significantly<br />

the sustained productivity and stability<br />

of various coniferous forest ecosystems<br />

within the Biome .<br />

5. To produce models of temporal and spatia l<br />

variations in coniferous forest ecosystem s<br />

or system components. These models will<br />

include factors affecting productivity an d<br />

stability of the systems and the linkages between<br />

terrestrial and aquatic environments ,<br />

forecasting the behavior of these systems<br />

and their relationships to human manipulation<br />

.<br />

6. To apply to specific models in the solutio n<br />

of major use problems in the Biome are a<br />

and assist other groups or agencies to d o<br />

likewise .<br />

Since these long-term objectives provid e<br />

only general guidance, considerable time an d<br />

effort is spent in defining annual researc h<br />

tasks in order to focus the research an d<br />

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