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PE EIE[R-Rg RESEARCH ON - HJ Andrews Experimental Forest

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where C is the percentage cover by epiphytes ,<br />

r is the radius at the base of the axis, and h i s<br />

the length of the axis . The bulk of epiphyte<br />

biomass for these branch systems is assume d<br />

to occur on the axes; if this is not the case ,<br />

the biomass of epiphytes on the many small<br />

branch systems, which are made up entirely<br />

of branchlets, may be significantly underestimated<br />

.<br />

Since EIV is based partly on subjective estimates<br />

of length of axes and percentage cove r<br />

by epiphytes, the relationship between EI V<br />

and epiphyte biomass may be expected t o<br />

vary from worker to worker and from sampling<br />

period to sampling period, and for thi s<br />

reason will be treated separately for each tree .<br />

However, it should be possible to correct<br />

importance values so that correlations may be<br />

obtained that will hold across a set of trees .<br />

Sampling Branch Systems<br />

The basic sampling unit for epiphytes o n<br />

axes in the branch systems is a "cylindrat, "<br />

which is analagous to a quadrat but runs completely<br />

around the axis so that two edges ar e<br />

fused . Our 1 dm cylindrats include the entir e<br />

surface for a distance of 1 dm along an axis .<br />

Thus the surface area of the cylindrat varies ,<br />

depending on the diameter of the axis in th e<br />

region sampled .<br />

Cylindrats were spaced along an axis with a<br />

distance of 4 dm from the center of on e<br />

cylindrat to the center of the next . Th e<br />

distance from the trunk to the first cylindra t<br />

sampled along a main axis was varied from 0<br />

to 1, 2, and 3 dm so as to avoid errors whic h<br />

would be introduced by horizontal zonation<br />

on the axis near its origin from the trunk .<br />

Estimates of epiphyte cover were made, an d<br />

then the epiphytes were stripped from th e<br />

cylindrat and bagged . Axis diameter at th e<br />

center of each cylindrat was measured t o<br />

enable calculation of surface area of the axes .<br />

Branchlets (

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