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PE EIE[R-Rg RESEARCH ON - HJ Andrews Experimental Forest

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asis of the structure of the different pin e<br />

crops (tables 1 and 5 and comparable descriptions<br />

from references in table 7) that the<br />

three lodgepole pine stands of this study carry<br />

appreciably greater burdens of respiratory<br />

biomass (Yoda et al. 1965). The major limitation<br />

of pine productivity in the Central and<br />

Northern Rocky Mountains may be the generally<br />

infertile soils and inimical continental<br />

climates. Under these environmental conditions<br />

not even the thinning treatment (stand<br />

4.3) and subsequent surge of productivity i n<br />

surviving stems gave increased yield on an are a<br />

basis (table 6) .<br />

Management decisions in the Colorad o<br />

Lodgepole Pine Zone must recognize th e<br />

inherent low productivity of P. contorta .<br />

Where wood production must continue t o<br />

have management priority, efficiency o f<br />

utilization can be increased at least 10 percent<br />

if live branches can be economically include d<br />

in the harvest (table 5) (Young 1968) . Management<br />

can also be directed toward uneven -<br />

aged stands for purposes of esthetics, enhanced<br />

groundcover production, or better<br />

game utilization. Small, irregularly shaped<br />

clearcut areas are also an attractive management<br />

possibility for long-term planning in this<br />

low productivity region . This study suggests<br />

strongly that because of its very low productivity,<br />

especially when compared against th e<br />

productivity of intensively managed pin e<br />

plantations, lodgepole pine stands in environments<br />

of site index of 90 or less should b e<br />

regarded as only minor resources of woo d<br />

harvest. A greater percentage of such land i n<br />

the lodgepole pine region of North Americ a<br />

might be devoted to recreational usage .<br />

A much neglected management tool in<br />

lodgepole pine forests is prescribed burning .<br />

Periodic fires were an important feature of<br />

lodgepole forests (Moir 1969) . This study and<br />

others (Kiil 1968) reveal that high quantities<br />

of slowly decomposable materials build up<br />

within natural stands . Fairly high quantities<br />

of nutrients become locked up in the forest<br />

floor humus (Moir and Grier 1969) ; this<br />

humus together with the shaded condition of<br />

the forest floor in closed stands has adverse<br />

effect upon the ground flora (Basile and<br />

Jensen 1971, Moir 1966). Controlled fire has<br />

at least four possible beneficial effects in<br />

lodgepole pine forests : (1) The reduction of<br />

fuel and wildfire probability, (2) stimulatio n<br />

of ground vegetation, (3) a nutrient "pulse"<br />

effect stimulating tree production, (4) tree<br />

thinning and conversion to uneven-age d<br />

stands. The possible use of controlled surfac e<br />

fires in P. contorta should be given<br />

considerably more attention .<br />

Acknowledgments<br />

Work reported in this paper was supporte d<br />

by the National Science Foundation, Grant No .<br />

B020357 in cooperation with the Coniferou s<br />

<strong>Forest</strong> Biome, U .S. Analysis of Ecosystems ,<br />

International Biological Program . This is Contribution<br />

No. 35 to the Coniferous <strong>Forest</strong><br />

Biome .<br />

Literature Cited<br />

Art, H. W., and P. L. Marks. 1971. A summary<br />

table of biomass and net annual primary<br />

production in forest ecosystems o f<br />

the world . In <strong>Forest</strong> biomass studies, 15th<br />

IUFRO Congr., p. 3-32 . Orono : Univ.<br />

Maine Press .<br />

Basile, J. V., and C. E. Jensen . 1971 . Grazing<br />

potential on lodgepole pine clearcuts i n<br />

Montana. USDA <strong>Forest</strong> Serv . Res. Pap.<br />

INT-98, 11 p . Intermountain <strong>Forest</strong> &<br />

Range Exp . Stn ., Ogden, Utah .<br />

Forrest, W. G. 1969. Variations in the accumulation,<br />

distribution and movement of<br />

mineral materials in radiata pine plantations.<br />

276 p. Ph .D. thesis on file at<br />

Australian Nat . Univ., Canberra .<br />

Hatiya, K., K. Doi, and R. Kobayashi. 1965 .<br />

Analysis of the growth in Japanese red pin e<br />

(Pinus densiflora) stands. A report on th e<br />

matured plantation in Iwate Prefecture .<br />

Govt. For. Exp. Stn. Bull. (Tokyo) 176 :<br />

75-88 .<br />

Johnstone, W . D. 1971 . Total standing cro p<br />

and tree component distributions in thre e<br />

stands of 100-year-old lodgepole pine . In<br />

<strong>Forest</strong> biomass studies, 15th IUFR O<br />

Congr., p. 81-89 . Orono : Univ. Maine Press .<br />

197

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