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longitudinally splitting and tattering post anthesis; flower cluster partially covered by a tubular 2-keeled prophyll and consisting of 5, 7, 9 or 11 flowers arranged in a group; 1-3 pistillate flowers in the centre subtended by 2 lateral cincinni of 0-2 pistillate and 2-4 staminate flowers, each flower, apart from the central pistillate, bearing an open, spathulate, 2-keeled, prophyllar bracteole. Staminate flowers symmetrical; calyx membranous, striate basally, stalked, tubular, with 3, short, triangular, apiculate lobes; corolla enclosed or only slightly exceeding the calyx, divided almost to the base into 3, elongate, striate, valvate petals; stamens 6, filaments united to form a thick fleshy, androecial tube, free from the corolla, tipped with 6 shallow lobes, bearing pendulous, rounded, latrorse anthers on the inside; pollen elliptic, monosulcate, with scabrate, tectate exine; pistillode very narrow, conical, slightly exceeding the androecial tube. Pistillate flowers superficially very similar to the staminate except slightly broader; calyx and corolla similar; staminodial tube bearing tiny empty anthers; gynoecium tricarpellate, triovulate, ± ellipsoidal, covered in reflexed scales, style long, narrow, 3angled; ovule basally attached, anatropous. Fruit ± spherical, stigmatic remains minute, conical; epicarp covered in vertical rows of rather thin reflexed scales, mesocarp very thin, almost obsolescent at maturity, endocarp not differentiated. Seed single, ± rounded, smooth or warty, basally attached with an oval hilum, covered with a thin, papery, sweet sarcotesta; endosperm homogenous, laterally penetrated by a smooth-margined mass of inner seed coat; embryo lateral opposite the intrusion. Germination adjacent-ligular; eophyll bifid, petiole of seedling dull reddish pink. Distribution Oncocalamus has a distinct Guinea-Congolian distribution and ranges from SE Nigeria to northern Angola, predominantly in coastal forest. Notes Recent collections have provided considerable insight into the life history and morphology of this intriguing genus. Although recent literature has stated that Oncocalamus is represented by only one, very variable, species, O. mannii, there are in fact four species present in Africa. A new species, O. tuleyi, previously assigned to O. mannii, is also described. 130
Key to the species of Oncocalamus Moderate to robust canes; stems with sheaths >20 mm in diameter; ocrea truncate with ± conspicuous 1.5-2.5 cm long rounded lobe; leaves, including cirrus, >1 m long, leaflets linear-lanceolate or only very mildly sigmoid, composed of a single fold: Stems with sheaths 30 mm in diameter, unarmed or moderately to profusely armed, with visible rounded swelling beneath leaf; leaf with >35 pairs of leaflets on each side of the rachis; inflorescence >1 m long; flower cluster with constant 3 pistillate flowers in each, seeds smooth: Leaf sheath moderately to profusely armed, ocrea almost horizontal, truncate, with small (8 cm long; rachillae deep crimson; north of the Sanaga River: O. tuleyi Slender canes; stems with sheaths
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THE TAXONOMY, ECOLOGY AND UTILISATI
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The final Chapter presents summaris
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CHAPTER TWO TAXONOMIC ACCOUNT 2.1.
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3.3.3.2. Climate 180 3.3.3.3. Topog
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CHAPTER SEVEN A SOCIO-ECONOMIC PROF
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REFERENCES 290 APPENDIX ONE INDIGEN
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Figure 42. Distribution of O. tuley
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Figure 110. How many years spent in
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ACKNOWLEDGEMENTS This work would no
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ecommended this project be funded b
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CHAPTER ONE MORPHOLOGY AND BIOGEOGR
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Similar branching of the stem is al
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The growing point of the stem of ra
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Aerial roots are commonly encounter
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sloughing of the sheath spines part
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For many of the species of Eremospa
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Figure 3. Acanthophylls of Laccospe
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In common with other members of the
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exception to this is Laccosperma op
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of overall diversity, when compared
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CHAPTER TWO TAXONOMIC ACCOUNT “Ta
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It is somewhat surpising that since
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evidence suggests that the climbing
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2.5. KEY TO THE GENERA Rattans clim
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horizontal, peduncle enclosed withi
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Knee absent: Stem ± triangular in
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15 cm broad, deeply notched with ro
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Distribution This species occurs fr
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Bot. Appl. 17: 896 (1936); Renier i
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Figure 3. Eremospatha cabrae (De Wi
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sterile, 1904 (BR!); Cabra s.n., Ma
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Figure 5. Eremospatha laurentii De
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(SCA!); de Wilde 2183, 60km S of Es
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long, decreasing distally, rachilla
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Distribution E. wendlandiana is dis
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indumentum; ocrea obliquely truncat
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Distribution E. barendii is known f
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stems sessile, up to 3.5 m long; ra
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Distribution E. macrocarpa is a ver
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Page 83 and 84: Distribution E. haullevilleana is r
Page 85 and 86: FHO!, BR!); Louis 9560, 20km W of Y
Page 87 and 88: order region of Cameroon and the Ri
Page 89 and 90: pairs c.3 cm long, at 45° angle to
Page 91 and 92: Figure 17. Eremospatha cuspidata (G
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Page 95 and 96: Figure 19. Eremospatha tessmanniana
Page 97 and 98: LACCOSPERMA (G. Mann & H. Wendl.) D
Page 99 and 100: Habitat and distribution The genus
Page 101 and 102: (1929); Hutch. in F.W.TA. 2: 391 (1
Page 103 and 104: Habitat and ecology Tolerant of dee
Page 105 and 106: Figure 21. Laccosperma opacum (G. M
Page 107 and 108: ± concolorous with prominent trans
Page 109 and 110: December 6, 1984 (WAG!); le Testu 1
Page 111 and 112: Figure 31. Laccosperma acutiflorum
Page 113 and 114: (06.09N:01.53W) Fl., June 1972 (MO!
Page 115 and 116: portion of stem; peduncle 12-20 cm
Page 117 and 118: Distribution L. robustum is very co
Page 119 and 120: 1997 (K!, EG!, BH!); Sunderland 179
Page 121 and 122: own indumentum below; leaflets comp
Page 123 and 124: Distribution This species is distri
Page 125 and 126: French). A brief discussion of this
Page 127 and 128: Imperfectly-known taxon Laccosperma
Page 129: ONCOCALAMUS (G. Mann & H. Wendl.) H
Page 133 and 134: moderately to sparsely armed with b
Page 135 and 136: Distribution O. mannii is restricte
Page 137 and 138: often concentrated on the sheath ap
Page 139 and 140: Distribution O. macrospathus is dis
Page 141 and 142: sheath, spines concentrated on marg
Page 143 and 144: Distribution This species is restri
Page 145 and 146: ocrea, often sloughing off to leave
Page 147 and 148: Notes O. wrightianus is distinct fr
Page 149 and 150: CALAMUS L. (Greek = a reed) L. in S
Page 151 and 152: Robyns & Tournay in Fl. du Parc Nat
Page 153 and 154: prickle-like spines; bracts tightly
Page 155 and 156: Distribution C. deërratus is the m
Page 157 and 158: (06.05N:00.50W) pist., March 17, 19
Page 159 and 160: MIXED COLLECTIONS A number of colle
Page 161 and 162: Figure 51. E. macrocarpa seedling,
Page 163 and 164: Figure 59. E. cuspidata, Etembue, E
Page 165 and 166: Figure 67. L. secundiflorum, Ghana
Page 167 and 168: Figure 75. O. mannii, Ayemaken, Equ
Page 169 and 170: CHAPTER THREE RATTAN DIVERSITY AND
Page 171 and 172: an inventory or survey. These param
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Page 175 and 176: 3.3.1.2 Climate The Campo Reserve h
Page 177 and 178: the reserve has no management plan
Page 179 and 180: associated with drier forest, was c
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the lowland forest is characterised
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sampling intensities were much lowe
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seedling 6 or length of stem) were
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Table 5. Rattan abundance and stock
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sampling of the rattan population,
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concentrated, not only on rattan, b
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Table 10. A comparison of rattan di
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4.1 INTRODUCTION CHAPTER FOUR RATTA
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the palm hearts from ground-level (
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feed on these fruits for long perio
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the genus Ceratogymna have been rec
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latter situation has been observed
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Hollow sheaths Some species of ant
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Table 15. Count of domatia and ratt
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Figure 85. Ant colonisation of leaf
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(Fisher et al., 1987), followed by
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Table 16. Hapaxanthy in the Palmae
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Figure 89. Lateral inflorescences o
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the absence of primate dispersers i
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CHAPTER SIX INDIGENOUS NOMENCLATURE
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The extensive nature and wide range
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Table 18. Summary of the non-cane u
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1977; Berlin, 1992). This hypothesi
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taxonomies are included in (or affi
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taxa (standing palms) within their
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Table 19. Life form, intermediate a
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For example, the Anyang of Cameroon
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Box 2. The structure of vernacular
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intermediate categories for African
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hoped that any future development o
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242
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CHAPTER SEVEN A SOCIO-ECONOMIC PROF
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Interestingly, the patterns of expl
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Figure 99. Scatterplot of size of m
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forest products (see Box 3) these a
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Figure 101. Mean range (or distance
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aid a wage. In fact, quite the oppo
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7.5.3 Socio-economic profile of the
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7.5.3.5 Previous occupations A larg
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the rainy season when transport dif
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Figure 114. Cited reasons for decli
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service. Unemployment increased fro
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Figure 116. Correlation between ran
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enefits rattan brings, those artisa
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from enabling the sustainable explo
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Figure 119. Woven basket products m
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such as Indonesia have lifted the b
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press; this study) and the conditio
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Table 31. Findings and recommendati
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of this trade, in fiscal terms, to
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Not threatened (species distributio
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clears the plot again to plant food
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forest products, such as rattan for
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from the informal forest economy an
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REFERENCES Abbiw, D. 1990. The usef
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Belcher, B. 1999. A production to c
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Burkill, I.H. 1935. A dictionary of
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Dalziel, J.M. 1937. The useful plan
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Dransfield, J. 1988b. The palms of
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Fisher, J.B. & J. Dransfield. 1977.
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Hall, J.B. & M.D. Swaine. 1981. Dis
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Huxley, C.R. 1978. The ant-plants M
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Lee, Y.F. 1993. Some models for est
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Moore, H.A. 1973. Palms in the trop
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Oteng-Amoako, A.A. & B. Obiri-Darko
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Randall, R. & E. Hunn. 1984. Do lif
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Siebert, S.F. 1997. Economically im
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Tenati, G. [in press]. The use of r
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Weiner, G. and Liese, W. 1989. Anat
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APPENDIX ONE INDIGENOUS NOMENCLATUR
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and Uganda, in the absence of large
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E. cuspidata (G. Mann & H. Wendl.)
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also a strong binding material (Rus
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(unpubl. notes): BENIN: Profizi (19
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L. laeve (G. Mann & H. Wendl.) H. W
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The “palm heart” is eaten widel
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A tea made from the young shoots is
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O. wrightianus Hutch. Vernacular na
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Bidgood & Vollesen: Tanzania; 3040
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Dransfield: Cameroon; 6998, 6999, 7
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Heudelot: Gambia; 372 Hoier: DR Con
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Lowe: Nigeria; 2792, 2793, 4353, Ca
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Reitsma: Gabon; 1340, 2047, 2151, 2
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Webb & Bullock: Cameroon; 310 Welle
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APPENDIX FOUR SOCIO-ECONOMIC SURVEY
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a) high forest b)farm c)fallow d) d
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APPENDIX FIVE PUBLICATIONS AND DISS
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DISSEMINATION 1. Sunderland, T.C.H.
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