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list of figures - Terry Sunderland

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CHAPTER ONE<br />

MORPHOLOGY AND BIOGEOGRAPHY OF AFRICAN RATTANS<br />

1.1 INTRODUCTION<br />

The rattans <strong>of</strong> Africa are represented by the endemic genera Laccosperma,<br />

Eremospatha and Oncocalamus, as well as by a single representative <strong>of</strong> the Asian<br />

genus Calamus. These climbing palms occur in a wide range ecological conditions<br />

within the lowland tropical forests <strong>of</strong> the continent and, throughout their range, play a<br />

significant role in the forest economy <strong>of</strong> the region through the utilisation <strong>of</strong> the inner<br />

stems, or cane. Despite this economic importance there has persisted considerable<br />

uncertainty with regard to the natural history <strong>of</strong> these species. This Chapter discusses<br />

the morphology and wider natural history <strong>of</strong> these species and provides the basis <strong>of</strong><br />

the taxonomic revision presented in Chapter 2.<br />

1.2 MORPHOLOGY<br />

1.2.1 Growth form<br />

All the rattan species <strong>of</strong> Africa are clustering i.e. they produce numerous stems from<br />

one individual. The suckers that develop to form a cluster are produced from the<br />

lowermost nodes <strong>of</strong> the original stem. They, in turn, produce further suckers and a<br />

large clump then develops (Dransfield, 1992a). Within the taxa <strong>of</strong> African rattans, the<br />

suckers are produced in the axils <strong>of</strong> the basal-most leaves and this is commonly the<br />

case with most clustering species <strong>of</strong> rattan. However, in a few instances (e.g. some<br />

species <strong>of</strong> the Asian genus Daemonorops), the sucker shoots are produced opposite to<br />

the leaf, whilst the vegetative buds <strong>of</strong> Korthalsia, another Asian genus, appear to be<br />

intermediate between axillary and leaf-opposed (Fisher and Dransfield, 1979).<br />

Although in general, rattan species are consistently single-stemmed or clustered it is<br />

possible to encounter individuals <strong>of</strong> predominantly clustering species with only a<br />

single stem e.g. the Asian species, Calamus subinermis H. Wendl ex Becc. This is<br />

sometimes the case with some normally clustering Eremospatha species in Africa,<br />

particularly in closed-canopy forest where the lack <strong>of</strong> light penetration could possibly<br />

be a limiting factor to the development <strong>of</strong> the cluster.<br />

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