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list of figures - Terry Sunderland

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exception to this is Laccosperma opacum, which possesses between 12-14 rows <strong>of</strong><br />

vertical scales.<br />

Within the outer covering <strong>of</strong> scales lies the remainder <strong>of</strong> the fruit wall. In the majority<br />

<strong>of</strong> the Calamoideae the fruit wall is rather thin (Uhl and Dransfield, 1987) but in the<br />

endemic African genera, there is a conspicuous mesocarp (this character is also shared<br />

by the genus Korthalsia (ibid.)), which is usually somewhat sweet-tasting. The<br />

innermost layer <strong>of</strong> the fruit wall, the pericarp, usually consists <strong>of</strong> a thin membrane<br />

within which lies the seed.<br />

A sarcotesta is a fleshy layer developed from the outer seed coat. This fleshy layer is<br />

common within the Asian members <strong>of</strong> the Calamoideae, notably among species where<br />

the fruit wall is thin and not particularly fleshy. Although rare within the African<br />

rattan taxa, the sarcotesta is particularly conspicuous in Calamus deërratus. Despite<br />

the presence <strong>of</strong> a fleshy mesocarp the seeds <strong>of</strong> the species <strong>of</strong> Oncocalamus also posses<br />

a very thin sarcotesta, as does Laccosperma opacum.<br />

1.2.21 The seed<br />

Usually within the Calamoideae, each fruit produces only one seed. However, some<br />

species <strong>of</strong> Laccosperma and Eremospatha sometimes have two, or perhaps three,<br />

seeds in each fruit, although this is rather uncommon. The species <strong>of</strong> both<br />

Laccosperma (with the exception <strong>of</strong> L. opacum, which is globose and has a deep cleft<br />

on one side) and Eremospatha have rounded seeds that are flattened on one side. The<br />

seeds <strong>of</strong> Oncocalamus are always globose. Those <strong>of</strong> Calamus deërratus are globose<br />

but possess a small conspicuous beak at the apex.<br />

The seed coat in a number <strong>of</strong> genera <strong>of</strong> African rattans shows some remarkable and<br />

wide variation. In the majority <strong>of</strong> the genus Laccosperma, the seed is smooth;<br />

however, Laccosperma opacum has a distinctly pitted and warty surface. Likewise, in<br />

Oncocalamus, the seed <strong>of</strong> O. mannii is also pitted, whilst the seed coat <strong>of</strong> the<br />

remaining species is smooth. The seeds <strong>of</strong> Eremospatha and Calamus are always<br />

smooth.<br />

37

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