altitude regions <strong>of</strong> east Africa >1500m. This species is usually found in forest under a canopy, but also occurs in open areas where it <strong>of</strong>ten forms dense thickets. Notes Beccari (1908) admitted the close relationship <strong>of</strong> the African species <strong>of</strong> Calamus he described, and his treatment <strong>of</strong> the African representatives <strong>of</strong> this genus, along those <strong>of</strong> Drude (1895) and de Wildeman (1904), was reasonable given the fragmentary and <strong>of</strong>ten rather poor quality material at their disposal. The fact that Calamus in Africa has been the cause <strong>of</strong> some taxonomic problems has been undoubtedly due to the recognition <strong>of</strong> poorly-defined infraspecific variation. However, from recent examination <strong>of</strong> herbarium specimens and field observations, it is clear that Calamus in Africa is represented by a single polymorphic species. Specimens examined GAMBIA: Anderson 131, Fr. July 10, 1975 (MO!); Heudelot 372, stam. 1839 (K!); Ingram s.n. sterile s.d. (K!); Starin 136, Aboko F.R., Fr., January 20, 1992 (K!); Starin 28, Aboko F.R., Fr., December 1979 (K!); SÈNÈGAL: Berhaut 877, Sangalkam (14.47N:17.12W) stam. December, 1950 (BR!); Vanden-Berghen 1752, Casamance (12.51N:15.17W) sterile, January 4, 1977 (MO!, BR!); Vanden- Berghen 4094, Badioure (12.53N:16.08W) stam. November 11, 1980 (BR!); Vanden-Berghen 5264, Bouyouye (12.26N:16.44W) pist. July 19, 1982 (BR!); GUINEA-CONAKRY: Chillou 1905, Kouyaya (10.25N:12.37W) sterile, March 19, 1940 (K!, BR!); Ory 216, Fula Kunda (13.15N:16.37W) sterile, January 25, 1954 (K!); SIERRA LEONE: Deighton 1847, Taisma, Central Province, Fr. July 8, 1930 (K!); Deighton 2592, Njala (08.06N:10.46W) sterile, January 1, 1933 (K!); Mann 895, Bagroo River (07.45N:12.50W) sterile, April 1861 (K!, FI!); Scott-Elliot 4738, Kambia (08.41N:13.03W) sterile, January 8, 1892 (K!); Scott-Elliot 5121, Musaia (09.27N:11.25W), sterile, March 10, 1892 (K!); Small 455, River Seli, Fr., September 5, 1951 (K!); Thomas 2753, Jigaya, stam. September 28, 1914 (K!); LIBERIA: Linder 1078, Piatah (07.12N:09.28W) sterile, October 15, 1926 (K!, MO!, WAG!); Linder 1116, Piatah (07.12N:09.28W) stam., October 17, 1926 (K!, MO!); Linder 1226, Gbanga (06.59N:09.28W) sterile, October 24, 1926 (K!, WAG!); Whyte s.n., near Kabatown (06.21N:10.43W) stam., April 1904 K!; CÔTE D'IVOIRE: Hepper & Maley 8041, Mont des Dans near Santa (08.16N:08.07W) pist., February 3, 1984 (K!); Hepper & Maley 8177, Taï Forest (05.38N:07.08W) sterile, February 9, 1984 (K!); Leeuwenberg 2524, 61km N <strong>of</strong> Sassandra (06.10N: 05.19W) stam., January 21, 1959 (WAG!); Leeuwenberg 2882, 18km NW <strong>of</strong> Sassandra (06.15N:05.00W) stam., November 26, 1960 (K!, WAG!); Oldeman 571, 9km ENE <strong>of</strong> Bereby (04.34N:07.00W) seedling, November 9, 1963 (WAG!); Oldeman 589, 3km E <strong>of</strong> Bereby (04.34N:07.00W) pist., November 5, 1963 (K!, WAG!); GHANA: Adams 2025, 2m E <strong>of</strong> Bibiani (06.20N:02.10W) sterile, December 23, 1953 (GC!); Cummins 128, Ashanti region, sterile, 1895 (K!); Enti 643, Kade Agricultural Station 156
(06.05N:00.50W) pist., March 17, 1972 (GC!, MO!, BR!); Enti & Hall s.n., Kade Agricultural Station (06.05N:00.50W) Fr., May 30, 1970 (GC!); Hall 2846, Anjakal, stam., January 20, 1965 (K!, GC!); Johnson 242, Kibbi-Akkim (06.09N:00.35W) Fr., December 13, 1899 (K!); Kinlock 3326, Tarkwa, Ndumnfri F.R. (05.10N:02.09W) sterile, February 15, 1934 (KUM!); Kisseadoo 435, Bobiri F.R. (06.38N:01.17W) sterile, November 10, 1988 (MO!); <strong>Sunderland</strong> 2262, Draw River Forest Reserve (05.12N:02.20W) sterile, May 26, 1999 (K!, KUM!); Tomlinson s.n., Bobiri F.R. (06.38N:01.17W) stam., December 20, 1957 (K!); Tomlinson s.n., Cape Coast F.R. (05.04N:01.30W) stam., December 15, 1957 (GC!); Tsiforkor s.n., Bunsu (06.15N:00.28W) sterile, November 22, 1995 (K!); Vigne 1868, Amentia F.R., E. Region (06.10N:01.58W) pist., March 30, 1930 (K!, KUM!); Vigne 3951, South Formango F.R. (06.35N:01.57W) sterile, July 30, 1935 (FHO!, KUM!); BENIN: Aufsess 424, Adjarra (06.32N:05.52E) sterile, December 6, 1988 (K!); NIGERIA: Allison 6994, Kabba Province (08.08N:06.44N) stam., November 15, 1943 (K!); Ayewoh 3854, Ondo Province, Ifon (06.54N:05.46E) sterile, February 24, 1944 (K!); Barter s.n., Onitsha (06.06N:06.48E) Fr., s.d. (K!); Bennett 8, sterile, February 24, 1906 (K!); Chapman 5010, Gangumi Forest Reserve, Gongola State (07.16N:11.23E) Fr., May 24, 1977 (K!); Chapman 5423, Baissa F.R., Gongola State (07.14N:10.38E) Fr., April 29, 1978 (K!); Gledhill 923, Akure F.R. (07.12N:05.11E) sterile, April 5, 1968 (K!, WAG!); Imp. Inst. Nigeria 347, Ahoada (05.03N:06.34E) sterile, February 1, 1936 (K!); Keay 28091, Kouton Kerifi, (08.08N:06.44N) Fl., November 6, 1950 (K!); Lowe 4353, Ilaro Forest Reserve, near Abeokuta (08.03N:06.06E) sterile, December 14, 1982 (K!); Tuley 846, Lagos (06.28N:03.20E) Fl., May 4, 1964 (K!); CAMEROON: unknown collector, Bezirk Djah (03.00N:12.40E) stam., March, 1910 (FI!); Buschen 3, Ebolowa (02.55N:11.08E) sterile, s.d. (FI!); Dransfield 6999, Mungo River Crossing (04.08N:09.31E) stam., June 27, 1991 (K!, SCA!); Dransfield 7000, Mungo River Crossing (04.08N:09.31E) sterile juvenile, June 27, 1991 (K!); Lederman 2428, Tibati (06.27N:12.37E) pist., January 29, 1909 (FI!); Mann 2147, Cameroon River (04.04N:09.38E) Fl. & Fr., January 1863 (K!, FI!); Meijer 15220, Sangmelima (02.55N:11.58E) sterile, March 24, 1981 (K!, MO!, WAG!, YA!); Meijer 15288, 9km W <strong>of</strong> Sangmelima (02.55N:11.58E) sterile, March 26, 1981 (WAG!); Mildbraed 4190, 21km northeast <strong>of</strong> Moloundou (02.03N:15.09E) Fl., June 6, 1911 (HBG!); Mildbraed 9548, Buea - Douala at Uham, Fl., May 5, 1914 (K!); Raynal 10150, 17km SE Ambam along the river Kye (02.23N:11.16E) sterile, March 1, 1963 (YA!); Raynal 10548, Guerima (7km NE de Bafia), gallery forest on river Mbam (04.44N:11.13E) stam., March 28, 1963 (YA!); <strong>Sunderland</strong> 1754, Limbe - Douala road at Mungo Bridge (04.08N:09.31E) stam., November 16, 1996 (K!, SCA!, BH!); <strong>Sunderland</strong> 1864, Djoum (02.48N:12.22E) sterile, September 9, 1997 (K!, YA!, WAG!); CENTRAL AFRICAN REPUBLIC: Fay 7020, N'Dele-Pata road (08.08N:21.08E) pist., May 30, 1985 (MO!); Fay 4381, Manovo-St Floris National Park (09.29N:21.17E) Fr., April 4, 1983 (K!); Harris & Fay 820, Ndakan (02.21N:16.09E) sterile, June 1, 1988 (K!); le Testu 3594, Haute-Kotto (04.11N:22.08E) Fr., January 17, 1922, (BR!); EQUATORIAL GUINEA: Tessmann 6, Fr., s.d. (FI!); GABON: Klaine 3246, Environs de Libreville (00.35N:09.22E) Fr., February 18, 1903 (K!); DR CONGO: Claessens 989, Mangbetu (02.28N:27.22E) sterile, June 1921 (BR!); de Witte 4066, Parc National de l'Upemba (09.04S:26.38E) pist., August, 1948 (BR!); Demeuse s.n., sterile, s.d. (BR!); Evrard 1876, Likimi (02.49N:20.44E) stam., September 29, 1955 (BR!); Evrard 3933, River Tshuapa en amont de Boende 157
- Page 1 and 2:
THE TAXONOMY, ECOLOGY AND UTILISATI
- Page 3 and 4:
The final Chapter presents summaris
- Page 5 and 6:
CHAPTER TWO TAXONOMIC ACCOUNT 2.1.
- Page 7 and 8:
3.3.3.2. Climate 180 3.3.3.3. Topog
- Page 9 and 10:
CHAPTER SEVEN A SOCIO-ECONOMIC PROF
- Page 11 and 12:
REFERENCES 290 APPENDIX ONE INDIGEN
- Page 13 and 14:
Figure 42. Distribution of O. tuley
- Page 15 and 16:
Figure 110. How many years spent in
- Page 17 and 18:
ACKNOWLEDGEMENTS This work would no
- Page 19 and 20:
ecommended this project be funded b
- Page 21 and 22:
CHAPTER ONE MORPHOLOGY AND BIOGEOGR
- Page 23 and 24:
Similar branching of the stem is al
- Page 25 and 26:
The growing point of the stem of ra
- Page 27 and 28:
Aerial roots are commonly encounter
- Page 29 and 30:
sloughing of the sheath spines part
- Page 31 and 32:
For many of the species of Eremospa
- Page 33 and 34:
Figure 3. Acanthophylls of Laccospe
- Page 35 and 36:
In common with other members of the
- Page 37 and 38:
exception to this is Laccosperma op
- Page 39 and 40:
of overall diversity, when compared
- Page 41 and 42:
CHAPTER TWO TAXONOMIC ACCOUNT “Ta
- Page 43 and 44:
It is somewhat surpising that since
- Page 45 and 46:
evidence suggests that the climbing
- Page 47 and 48:
2.5. KEY TO THE GENERA Rattans clim
- Page 49 and 50:
horizontal, peduncle enclosed withi
- Page 51 and 52:
Knee absent: Stem ± triangular in
- Page 53 and 54:
15 cm broad, deeply notched with ro
- Page 55 and 56:
Distribution This species occurs fr
- Page 57 and 58:
Bot. Appl. 17: 896 (1936); Renier i
- Page 59 and 60:
Figure 3. Eremospatha cabrae (De Wi
- Page 61 and 62:
sterile, 1904 (BR!); Cabra s.n., Ma
- Page 63 and 64:
Figure 5. Eremospatha laurentii De
- Page 65 and 66:
(SCA!); de Wilde 2183, 60km S of Es
- Page 67 and 68:
long, decreasing distally, rachilla
- Page 69 and 70:
Distribution E. wendlandiana is dis
- Page 71 and 72:
indumentum; ocrea obliquely truncat
- Page 73 and 74:
Distribution E. barendii is known f
- Page 75 and 76:
stems sessile, up to 3.5 m long; ra
- Page 77 and 78:
Distribution E. macrocarpa is a ver
- Page 79 and 80:
(YA!); Letouzey 12563, Lac Tissongo
- Page 81 and 82:
and sparse distally; leaflets 8-14
- Page 83 and 84:
Distribution E. haullevilleana is r
- Page 85 and 86:
FHO!, BR!); Louis 9560, 20km W of Y
- Page 87 and 88:
order region of Cameroon and the Ri
- Page 89 and 90:
pairs c.3 cm long, at 45° angle to
- Page 91 and 92:
Figure 17. Eremospatha cuspidata (G
- Page 93 and 94:
ase, entire and acuminate to irregu
- Page 95 and 96:
Figure 19. Eremospatha tessmanniana
- Page 97 and 98:
LACCOSPERMA (G. Mann & H. Wendl.) D
- Page 99 and 100:
Habitat and distribution The genus
- Page 101 and 102:
(1929); Hutch. in F.W.TA. 2: 391 (1
- Page 103 and 104:
Habitat and ecology Tolerant of dee
- Page 105 and 106: Figure 21. Laccosperma opacum (G. M
- Page 107 and 108: ± concolorous with prominent trans
- Page 109 and 110: December 6, 1984 (WAG!); le Testu 1
- Page 111 and 112: Figure 31. Laccosperma acutiflorum
- Page 113 and 114: (06.09N:01.53W) Fl., June 1972 (MO!
- Page 115 and 116: portion of stem; peduncle 12-20 cm
- Page 117 and 118: Distribution L. robustum is very co
- Page 119 and 120: 1997 (K!, EG!, BH!); Sunderland 179
- Page 121 and 122: own indumentum below; leaflets comp
- Page 123 and 124: Distribution This species is distri
- Page 125 and 126: French). A brief discussion of this
- Page 127 and 128: Imperfectly-known taxon Laccosperma
- Page 129 and 130: ONCOCALAMUS (G. Mann & H. Wendl.) H
- Page 131 and 132: Key to the species of Oncocalamus M
- Page 133 and 134: moderately to sparsely armed with b
- Page 135 and 136: Distribution O. mannii is restricte
- Page 137 and 138: often concentrated on the sheath ap
- Page 139 and 140: Distribution O. macrospathus is dis
- Page 141 and 142: sheath, spines concentrated on marg
- Page 143 and 144: Distribution This species is restri
- Page 145 and 146: ocrea, often sloughing off to leave
- Page 147 and 148: Notes O. wrightianus is distinct fr
- Page 149 and 150: CALAMUS L. (Greek = a reed) L. in S
- Page 151 and 152: Robyns & Tournay in Fl. du Parc Nat
- Page 153 and 154: prickle-like spines; bracts tightly
- Page 155: Distribution C. deërratus is the m
- Page 159 and 160: MIXED COLLECTIONS A number of colle
- Page 161 and 162: Figure 51. E. macrocarpa seedling,
- Page 163 and 164: Figure 59. E. cuspidata, Etembue, E
- Page 165 and 166: Figure 67. L. secundiflorum, Ghana
- Page 167 and 168: Figure 75. O. mannii, Ayemaken, Equ
- Page 169 and 170: CHAPTER THREE RATTAN DIVERSITY AND
- Page 171 and 172: an inventory or survey. These param
- Page 173 and 174: clustering species are to be includ
- Page 175 and 176: 3.3.1.2 Climate The Campo Reserve h
- Page 177 and 178: the reserve has no management plan
- Page 179 and 180: associated with drier forest, was c
- Page 181 and 182: the lowland forest is characterised
- Page 183 and 184: sampling intensities were much lowe
- Page 185 and 186: seedling 6 or length of stem) were
- Page 187 and 188: Table 5. Rattan abundance and stock
- Page 189 and 190: sampling of the rattan population,
- Page 191 and 192: concentrated, not only on rattan, b
- Page 193 and 194: Table 10. A comparison of rattan di
- Page 195 and 196: 4.1 INTRODUCTION CHAPTER FOUR RATTA
- Page 197 and 198: the palm hearts from ground-level (
- Page 199 and 200: feed on these fruits for long perio
- Page 201 and 202: the genus Ceratogymna have been rec
- Page 203 and 204: latter situation has been observed
- Page 205 and 206: Hollow sheaths Some species of ant
- Page 207 and 208:
Table 15. Count of domatia and ratt
- Page 209 and 210:
Figure 85. Ant colonisation of leaf
- Page 211 and 212:
(Fisher et al., 1987), followed by
- Page 213 and 214:
Table 16. Hapaxanthy in the Palmae
- Page 215 and 216:
Figure 89. Lateral inflorescences o
- Page 217 and 218:
the absence of primate dispersers i
- Page 219 and 220:
CHAPTER SIX INDIGENOUS NOMENCLATURE
- Page 221:
The extensive nature and wide range
- Page 224 and 225:
Table 18. Summary of the non-cane u
- Page 226 and 227:
1977; Berlin, 1992). This hypothesi
- Page 228 and 229:
taxonomies are included in (or affi
- Page 230 and 231:
taxa (standing palms) within their
- Page 232 and 233:
Table 19. Life form, intermediate a
- Page 234 and 235:
For example, the Anyang of Cameroon
- Page 236 and 237:
Box 2. The structure of vernacular
- Page 238 and 239:
intermediate categories for African
- Page 240 and 241:
hoped that any future development o
- Page 242 and 243:
242
- Page 244 and 245:
CHAPTER SEVEN A SOCIO-ECONOMIC PROF
- Page 246 and 247:
Interestingly, the patterns of expl
- Page 248 and 249:
Figure 99. Scatterplot of size of m
- Page 250 and 251:
forest products (see Box 3) these a
- Page 252 and 253:
Figure 101. Mean range (or distance
- Page 254 and 255:
aid a wage. In fact, quite the oppo
- Page 256 and 257:
7.5.3 Socio-economic profile of the
- Page 258 and 259:
7.5.3.5 Previous occupations A larg
- Page 260 and 261:
the rainy season when transport dif
- Page 262 and 263:
Figure 114. Cited reasons for decli
- Page 264 and 265:
service. Unemployment increased fro
- Page 266 and 267:
Figure 116. Correlation between ran
- Page 268 and 269:
enefits rattan brings, those artisa
- Page 270 and 271:
from enabling the sustainable explo
- Page 272 and 273:
Figure 119. Woven basket products m
- Page 274 and 275:
such as Indonesia have lifted the b
- Page 276 and 277:
press; this study) and the conditio
- Page 278 and 279:
Table 31. Findings and recommendati
- Page 280 and 281:
of this trade, in fiscal terms, to
- Page 282 and 283:
Not threatened (species distributio
- Page 284 and 285:
clears the plot again to plant food
- Page 286 and 287:
forest products, such as rattan for
- Page 288 and 289:
from the informal forest economy an
- Page 290 and 291:
REFERENCES Abbiw, D. 1990. The usef
- Page 292 and 293:
Belcher, B. 1999. A production to c
- Page 294 and 295:
Burkill, I.H. 1935. A dictionary of
- Page 296 and 297:
Dalziel, J.M. 1937. The useful plan
- Page 298 and 299:
Dransfield, J. 1988b. The palms of
- Page 300 and 301:
Fisher, J.B. & J. Dransfield. 1977.
- Page 302 and 303:
Hall, J.B. & M.D. Swaine. 1981. Dis
- Page 304 and 305:
Huxley, C.R. 1978. The ant-plants M
- Page 306 and 307:
Lee, Y.F. 1993. Some models for est
- Page 308 and 309:
Moore, H.A. 1973. Palms in the trop
- Page 310 and 311:
Oteng-Amoako, A.A. & B. Obiri-Darko
- Page 312 and 313:
Randall, R. & E. Hunn. 1984. Do lif
- Page 314 and 315:
Siebert, S.F. 1997. Economically im
- Page 316 and 317:
Tenati, G. [in press]. The use of r
- Page 318 and 319:
Weiner, G. and Liese, W. 1989. Anat
- Page 320 and 321:
APPENDIX ONE INDIGENOUS NOMENCLATUR
- Page 322 and 323:
and Uganda, in the absence of large
- Page 324 and 325:
E. cuspidata (G. Mann & H. Wendl.)
- Page 326 and 327:
also a strong binding material (Rus
- Page 328 and 329:
(unpubl. notes): BENIN: Profizi (19
- Page 330 and 331:
L. laeve (G. Mann & H. Wendl.) H. W
- Page 332 and 333:
The “palm heart” is eaten widel
- Page 334 and 335:
A tea made from the young shoots is
- Page 336 and 337:
O. wrightianus Hutch. Vernacular na
- Page 338 and 339:
Bidgood & Vollesen: Tanzania; 3040
- Page 340 and 341:
Dransfield: Cameroon; 6998, 6999, 7
- Page 342 and 343:
Heudelot: Gambia; 372 Hoier: DR Con
- Page 344 and 345:
Lowe: Nigeria; 2792, 2793, 4353, Ca
- Page 346 and 347:
Reitsma: Gabon; 1340, 2047, 2151, 2
- Page 348 and 349:
Webb & Bullock: Cameroon; 310 Welle
- Page 350 and 351:
APPENDIX FOUR SOCIO-ECONOMIC SURVEY
- Page 352 and 353:
a) high forest b)farm c)fallow d) d
- Page 354 and 355:
APPENDIX FIVE PUBLICATIONS AND DISS
- Page 356 and 357:
DISSEMINATION 1. Sunderland, T.C.H.