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located the district offices and the only hospital and police station in the South<br />

Rupununi. A secondary school was opened in this village in 1999, but although the<br />

provision of secondary education in the area is welcomed the headmaster has<br />

complained about a chronic shortage of appropriately trained staff. Karaudanawa to<br />

the west is situated on the upper reaches of the Rupununi river, and Achiwuib lies still<br />

further west, its reservation boundary abutting the Brazilian border. Several smaller<br />

settlements are administered by these villages, and villagers of both Achiwuib and<br />

Karaudanawa have established separate farming settlements to the south of the<br />

villages, close to the forest-savannah boundary, as the farming areas are at<br />

considerable distances from the villages themselves.<br />

Wapishana settlement in Guyana thus encompasses almost the entire forest-<br />

savannah boundary in the South Rupununi, and the range of habitat types found<br />

within this. Analysis of changes in the forest-savannah boundary over the period<br />

1952-1983 shows it to have remained stable over time, despite increases in<br />

population and permanence of settlements and hence apparent agricultural<br />

intensification. <strong>In</strong> fact, in one uncultivated area north of Shea the forest appears to<br />

be advancing into the savannah (Eden 1986). Preliminary research findings based on<br />

a study of genetic variation of bird species across savannah-forest ecotones in<br />

Cameroon suggest that such transitional habitats may be of great importance in<br />

generating biodiversity over evolutionary time-scales, via the promotion of<br />

intraspecific genetic diversity and hence speciation events (Smith et al. 1997).<br />

Should these findings be confirmed, and should they also apply to such habitats in<br />

South America, this would have great implications for the conservation importance of<br />

the area occupied by the Wapishana in Guyana. As a major part of an ecotone of the<br />

world’s largest intact area of tropical forest, it would represent an area whose<br />

biodiversity is of global importance. Human use of and effects upon this habitat is<br />

thus a subject of key interest to those concerned with global conservation, and<br />

comprises the subject matter of the next chapter.<br />

3.2.4 Maruranau as an example of a modern Wapishana village<br />

The major site of field research in this study was Maruranau village, which was<br />

founded in 1922 after converts of Fr. Cary-Elwes resident in the village of<br />

Sawaramanirnao decided to relocate to a previously uninhabited site on the banks of<br />

Marora wa'o creek, owing to a shortage of water at the former location. Work started<br />

on the church in 1922, and Cary-Elwes reported the village to have relocated<br />

following its completion by the time of his visit in 1923 (Bridges 1985: 146, 158,

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