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Culture and Ecology of Chaco Canyon and the San Juan Basin

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Environment <strong>and</strong> Natural Resources 41I'SS: Sarcobatus vermiculatus-Sporobolus airoides.At <strong>the</strong> eastern end <strong>of</strong> <strong>the</strong> park, from where <strong>the</strong><strong>Chaco</strong> <strong>and</strong> Gallo washes cross <strong>the</strong> boundary to <strong>the</strong>road that crosses <strong>the</strong> <strong>Chaco</strong> Wash from <strong>the</strong> south exitto <strong>the</strong> visitor center, are st<strong>and</strong>s <strong>of</strong> black greasewood(Sarcobatus vermiculatus). Here <strong>the</strong> soils are characteristicallyalkaline silt <strong>and</strong> clay. Also in this areaare rubber rabbitbrush (Chrysothamnus nauseosus),several species <strong>of</strong> wheatgrasses (Agropyron), alkalisacaton (Sporobolus airoides), <strong>and</strong> spiny or shadscalesaltbush (Atriplex confertifolia).PJ: Pinon-juniper. This woodl<strong>and</strong> zone, foundonly on Chacra Mesa at <strong>the</strong> sou<strong>the</strong>astern edge <strong>of</strong> <strong>the</strong>park, is dominated by one-seed juniper (Juniperusmonosperma), with pinon (Pinus edulis) below 2,133m (7,000 ft). Shrubs include true mountain mahogany(Cerocarpus montanus) <strong>and</strong> black sagebrush(Artemesia nova), plus a large variety <strong>of</strong> grasses <strong>and</strong>forbs that have no indicator value.RW: Riparian woody vegetation. The entirestream <strong>of</strong> <strong>the</strong> <strong>Chaco</strong> Wash was characterized as riparian,<strong>and</strong> woody, to correspond with soil piping <strong>and</strong>erosional studies being conducted by Malde. Severalspecies <strong>of</strong> rabbitbrush (Chrysothamnus spp.), greasewood(Sarcobatus sp.), coyote willow (Salix exigua),four-wing saltbush (Atriplex canescens), Fremont'scottonwood (Populus fremontii), <strong>and</strong> <strong>the</strong> exotictamarisk (Tamarix pent<strong>and</strong>ra) are present.D: Dune area. Da: dune area-active. Dv: dunearea-vegetated. Rubber rabbitbrush (Chrysothamnusnauseosus) <strong>and</strong> s<strong>and</strong> sagebrush (Artemisiafilifolia) are<strong>the</strong> dominant species present. Also present are severalherbs <strong>and</strong> grasses.BR: Bare rock.Wt: Watershed treatment. Manmade zone.Of: Old field succession. Manmade zone.Potter <strong>and</strong> Kelley (1980: 103) indicated thatchanges in vegetation could be linked to relativelyminor changes in environmental conditions. Manyspecies in <strong>the</strong> area, especially <strong>the</strong> annuals, fluctuategreatly in density <strong>and</strong> coverage due to changes inannual precipitation. There is also a close link t<strong>of</strong>eatures <strong>of</strong> geology, physiography, <strong>and</strong> soils. Insummary, a coarse vegetative classification map, witha detailed description <strong>of</strong> <strong>the</strong> different vegetative zones,was completed, <strong>and</strong> reasons for some changes invegetation were noted. A. Cully (1985a:Appendix A)published a comprehensive plant list based on existingcollections in <strong>the</strong> park <strong>and</strong> supplemented by thosefrom Ron Bronitsky in 1972 <strong>and</strong> those that she <strong>and</strong>Mollie Struever Toll collected in association withpollen <strong>and</strong> flotation studies during <strong>the</strong> <strong>Chaco</strong> Project.After <strong>the</strong> monument became a park in 1980, <strong>the</strong>additional l<strong>and</strong>s were surveyed by Floyd-Hanna <strong>and</strong>Hanna (1995) <strong>and</strong> Floyd-Hanna et al. (1993). Thevegetative map by Kelley <strong>and</strong> Potter was digitized,<strong>and</strong> inventory transects provided data, indicating that13 distinct vegetation types can be defined <strong>and</strong>aggregated into eight visually distinct vegetationcommunities. These include <strong>the</strong> riparian wash,Artemesia tridetatalChrysothamnus nauseosus, Sarcobatusassociations, pinon/juniper associations, Artemesianova-J.monosperma-C. visiiflorus, A. nova­Gutierrezia sarothrae-C. greenei, Sporobolus airoidesassociations, <strong>and</strong> Hilaria jamesii-Sarcobatus vermiculatus-Boutelouagracilis-Orozopsis hymenoides, aswell as <strong>the</strong> distinct formations <strong>of</strong> s<strong>and</strong>stone cliffs <strong>and</strong>active dunes.Scott (1980) confirmed <strong>the</strong> variability in plantresponse to available moisture <strong>and</strong> supported observations<strong>and</strong> explanations proposed by Potter (1974)regarding <strong>the</strong> annual variability in plant species due todifferences in rainfall. From 1975 through 1977,rainfall in <strong>Chaco</strong> was below normal (215 mm). In1975, it was 68 percent <strong>of</strong> average; in 1976, 62percent; <strong>and</strong> in 1977, 87 percent. When monthlyobservations on <strong>the</strong> phenology <strong>of</strong> plants in samplingareas were compared, seven <strong>of</strong> 11 observed species <strong>of</strong>perennial plants reproduced during 1976, while 10 <strong>of</strong>11 reproduced in 1977. The flowering period (Aprilto October) contained two peaks: May-June, <strong>and</strong>August-October. Norman <strong>and</strong> Duke (section 2 <strong>of</strong>"Phenology <strong>of</strong> <strong>the</strong> Perennial Plants") recorded differingflowering dates for various species, some <strong>of</strong> whichflowered twice; e.g., gall eta grass (Hilaria james;;)<strong>and</strong> muhly (Muhlenbergia sp.). Scott (section 7 on"Perennial Plant Coverage") noted that some perennialspartially or completely died (e.g., snakeweed[Gutierrezia sp.]), depending on available moisture,while o<strong>the</strong>rs (e. g. , Indian ricegrass [Oryzopsishymenoides], ephedra [Ephedra torreyi]) , blue gramagrass [Boutelous gracilis J, cactus [Opuntia polya-

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