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130925-studie-wildlife-comeback-in-europe

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Scale Status Population trend Justification Threats<br />

Global<br />

[2]<br />

Least Concern Decreas<strong>in</strong>g Widespread distribution<br />

Large population size<br />

1. Residential & commercial development<br />

2. Agriculture & aquaculture (livestock farm<strong>in</strong>g)<br />

3. Transportation and service corridors<br />

4. Hunt<strong>in</strong>g & persecution<br />

5. Human disturbance<br />

Europe<br />

[12]<br />

Vulnerable Decreas<strong>in</strong>g Small population size<br />

Decl<strong>in</strong>es outside the northern part of<br />

the range<br />

1. Residential & commercial development<br />

2. Agriculture & aquaculture (livestock farm<strong>in</strong>g)<br />

3. Hunt<strong>in</strong>g & persecution<br />

4. Human disturbance<br />

Europe –<br />

regional<br />

population<br />

Vulnerable:<br />

Sweden [23]<br />

Endangered:<br />

Norway [24] ,<br />

Karelia [12] N/A Scand<strong>in</strong>avia [12] : Stable but small<br />

population, limited connectivity,<br />

genetically dist<strong>in</strong>ct<br />

Karelia [12] : Small and decl<strong>in</strong><strong>in</strong>g<br />

Scand<strong>in</strong>avia [12] : Hunt<strong>in</strong>g & persecution<br />

Karelia [12] : Poach<strong>in</strong>g, Depletion of prey base<br />

Habitat preferences and general densities<br />

In Europe, the Wolver<strong>in</strong>e has a broad range [1] ,<br />

occurr<strong>in</strong>g primarily <strong>in</strong> boreal forest and tundra [4] .<br />

Because Wolver<strong>in</strong>es exist at low densities [4] and<br />

have large home ranges, they require vast areas of<br />

suitable habitat for viable breed<strong>in</strong>g populations [1] .<br />

Males have significantly larger home ranges, which<br />

overlap with females but are exclusive with other<br />

males [13] . Den sites are placed primarily <strong>in</strong> steep,<br />

rugged terra<strong>in</strong> with bare rock, which is some<br />

distance from <strong>in</strong>frastructure such as roads [14] .<br />

Wolver<strong>in</strong>es select habitats that promote survival<br />

through limited encounters with humans but<br />

which are rich <strong>in</strong> prey, and this selection is stronger<br />

<strong>in</strong> w<strong>in</strong>ter [15] . The species occurs at extremely low<br />

densities of between 0.1 and 1.5 <strong>in</strong>dividuals per<br />

100 km 2 [1] .<br />

Legal protection and conservation status<br />

The Wolver<strong>in</strong>e is protected under the Habitats<br />

Directive (Annexes II and IV) <strong>in</strong> Sweden and<br />

F<strong>in</strong>land [16] and the Bern Convention (Appendix<br />

II) <strong>in</strong> all three Scand<strong>in</strong>avian countries [17] . Limited<br />

culls are undertaken <strong>in</strong> Sweden, and Norway<br />

heavily manages the species to control numbers [18] .<br />

Compensations schemes are <strong>in</strong> place for damages<br />

to semi-domesticated re<strong>in</strong>deer (all three countries)<br />

and sheep (Norway and Sweden) [18, 19] . Much of the<br />

biology and ecology of the Wolver<strong>in</strong>e was not<br />

known until fairly recently, but highly cooperative<br />

research projects are now <strong>in</strong> place <strong>in</strong> Sweden [20] ,<br />

F<strong>in</strong>land [21] , and Norway [18] to collect miss<strong>in</strong>g <strong>in</strong>formation<br />

about wolver<strong>in</strong>e ecology, for <strong>in</strong>stance<br />

assess<strong>in</strong>g the impact of the species on re<strong>in</strong>deer,<br />

and explor<strong>in</strong>g <strong>in</strong>teractions with the Eurasian lynx<br />

(Lynx lynx) [20, 22] .<br />

At a global level, the Wolver<strong>in</strong>e is classified as<br />

Least Concern due to the species’ wide distribution<br />

and large population size, as well as on-go<strong>in</strong>g<br />

expansion <strong>in</strong>to its former range (Table 1). However,<br />

densities rema<strong>in</strong> low, and the overall species trend<br />

is believed to be decl<strong>in</strong><strong>in</strong>g <strong>in</strong> number despite<br />

signs of range recovery (Table 1). In Europe, the<br />

species is currently classified as Vulnerable with a<br />

decreas<strong>in</strong>g population trend (Table 1). Despite this,<br />

extensive hunt<strong>in</strong>g quotas and lethal control, which<br />

are <strong>in</strong>directly related to depredation conflicts, are<br />

implemented <strong>in</strong> Norway to decrease population<br />

size, while Sweden only culls the species at a very<br />

local level [4, 5] .<br />

Abundance and distribution:<br />

current status<br />

Recent population estimates suggest that the<br />

current European population comprises just over<br />

2,600 <strong>in</strong>dividuals (Table 2), form<strong>in</strong>g two populations<br />

between which there is believed to be some<br />

exchange [18] : the Karelian population (European<br />

Russia and eastern F<strong>in</strong>land) and the Scand<strong>in</strong>avian<br />

population (Sweden, Norway and the rest of<br />

F<strong>in</strong>land). At a country level, the largest populations<br />

occur <strong>in</strong> Russia and Sweden, which account for 53%<br />

and 26% of the European population respectively<br />

(Table 2). Smaller populations exist <strong>in</strong> Norway (15%)<br />

and F<strong>in</strong>land (6%). It should be noted, however, that<br />

the estimate for European Russia may be outdated.<br />

Of the largest population of the species <strong>in</strong><br />

Russia (based on <strong>in</strong>formation from 2005), the<br />

majority occurred <strong>in</strong> the Komi Republic accord<strong>in</strong>g<br />

to a pre-2000 estimate [1] . Although undoubtedly<br />

important <strong>in</strong> terms of size, the Russian population<br />

Table 1.<br />

Summary of Global<br />

and European Red<br />

List assessments and<br />

threats listed for the<br />

Wolver<strong>in</strong>e.<br />

Table 2.<br />

Latest approximate<br />

population<br />

estimates for the<br />

Wolver<strong>in</strong>e <strong>in</strong> Europe<br />

and for European<br />

populations.<br />

Please note that<br />

the estimate for<br />

European Russia may<br />

be outdated.<br />

Estimate Year assessed Reference<br />

Global No data -<br />

[2]<br />

Europe 2,630–2,640 2005–11<br />

[11, 18]<br />

% of global population No data<br />

European Russia 1,400 2005<br />

[11]<br />

F<strong>in</strong>land 165–175 2011<br />

[18]<br />

Norway 385 2011<br />

[18]<br />

Sweden 680 2011<br />

[18]<br />

121

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