130925-studie-wildlife-comeback-in-europe
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Scale Status Population trend Justification Threats<br />
Global<br />
[2]<br />
Least Concern Decreas<strong>in</strong>g Widespread distribution<br />
Large population size<br />
1. Residential & commercial development<br />
2. Agriculture & aquaculture (livestock farm<strong>in</strong>g)<br />
3. Transportation and service corridors<br />
4. Hunt<strong>in</strong>g & persecution<br />
5. Human disturbance<br />
Europe<br />
[12]<br />
Vulnerable Decreas<strong>in</strong>g Small population size<br />
Decl<strong>in</strong>es outside the northern part of<br />
the range<br />
1. Residential & commercial development<br />
2. Agriculture & aquaculture (livestock farm<strong>in</strong>g)<br />
3. Hunt<strong>in</strong>g & persecution<br />
4. Human disturbance<br />
Europe –<br />
regional<br />
population<br />
Vulnerable:<br />
Sweden [23]<br />
Endangered:<br />
Norway [24] ,<br />
Karelia [12] N/A Scand<strong>in</strong>avia [12] : Stable but small<br />
population, limited connectivity,<br />
genetically dist<strong>in</strong>ct<br />
Karelia [12] : Small and decl<strong>in</strong><strong>in</strong>g<br />
Scand<strong>in</strong>avia [12] : Hunt<strong>in</strong>g & persecution<br />
Karelia [12] : Poach<strong>in</strong>g, Depletion of prey base<br />
Habitat preferences and general densities<br />
In Europe, the Wolver<strong>in</strong>e has a broad range [1] ,<br />
occurr<strong>in</strong>g primarily <strong>in</strong> boreal forest and tundra [4] .<br />
Because Wolver<strong>in</strong>es exist at low densities [4] and<br />
have large home ranges, they require vast areas of<br />
suitable habitat for viable breed<strong>in</strong>g populations [1] .<br />
Males have significantly larger home ranges, which<br />
overlap with females but are exclusive with other<br />
males [13] . Den sites are placed primarily <strong>in</strong> steep,<br />
rugged terra<strong>in</strong> with bare rock, which is some<br />
distance from <strong>in</strong>frastructure such as roads [14] .<br />
Wolver<strong>in</strong>es select habitats that promote survival<br />
through limited encounters with humans but<br />
which are rich <strong>in</strong> prey, and this selection is stronger<br />
<strong>in</strong> w<strong>in</strong>ter [15] . The species occurs at extremely low<br />
densities of between 0.1 and 1.5 <strong>in</strong>dividuals per<br />
100 km 2 [1] .<br />
Legal protection and conservation status<br />
The Wolver<strong>in</strong>e is protected under the Habitats<br />
Directive (Annexes II and IV) <strong>in</strong> Sweden and<br />
F<strong>in</strong>land [16] and the Bern Convention (Appendix<br />
II) <strong>in</strong> all three Scand<strong>in</strong>avian countries [17] . Limited<br />
culls are undertaken <strong>in</strong> Sweden, and Norway<br />
heavily manages the species to control numbers [18] .<br />
Compensations schemes are <strong>in</strong> place for damages<br />
to semi-domesticated re<strong>in</strong>deer (all three countries)<br />
and sheep (Norway and Sweden) [18, 19] . Much of the<br />
biology and ecology of the Wolver<strong>in</strong>e was not<br />
known until fairly recently, but highly cooperative<br />
research projects are now <strong>in</strong> place <strong>in</strong> Sweden [20] ,<br />
F<strong>in</strong>land [21] , and Norway [18] to collect miss<strong>in</strong>g <strong>in</strong>formation<br />
about wolver<strong>in</strong>e ecology, for <strong>in</strong>stance<br />
assess<strong>in</strong>g the impact of the species on re<strong>in</strong>deer,<br />
and explor<strong>in</strong>g <strong>in</strong>teractions with the Eurasian lynx<br />
(Lynx lynx) [20, 22] .<br />
At a global level, the Wolver<strong>in</strong>e is classified as<br />
Least Concern due to the species’ wide distribution<br />
and large population size, as well as on-go<strong>in</strong>g<br />
expansion <strong>in</strong>to its former range (Table 1). However,<br />
densities rema<strong>in</strong> low, and the overall species trend<br />
is believed to be decl<strong>in</strong><strong>in</strong>g <strong>in</strong> number despite<br />
signs of range recovery (Table 1). In Europe, the<br />
species is currently classified as Vulnerable with a<br />
decreas<strong>in</strong>g population trend (Table 1). Despite this,<br />
extensive hunt<strong>in</strong>g quotas and lethal control, which<br />
are <strong>in</strong>directly related to depredation conflicts, are<br />
implemented <strong>in</strong> Norway to decrease population<br />
size, while Sweden only culls the species at a very<br />
local level [4, 5] .<br />
Abundance and distribution:<br />
current status<br />
Recent population estimates suggest that the<br />
current European population comprises just over<br />
2,600 <strong>in</strong>dividuals (Table 2), form<strong>in</strong>g two populations<br />
between which there is believed to be some<br />
exchange [18] : the Karelian population (European<br />
Russia and eastern F<strong>in</strong>land) and the Scand<strong>in</strong>avian<br />
population (Sweden, Norway and the rest of<br />
F<strong>in</strong>land). At a country level, the largest populations<br />
occur <strong>in</strong> Russia and Sweden, which account for 53%<br />
and 26% of the European population respectively<br />
(Table 2). Smaller populations exist <strong>in</strong> Norway (15%)<br />
and F<strong>in</strong>land (6%). It should be noted, however, that<br />
the estimate for European Russia may be outdated.<br />
Of the largest population of the species <strong>in</strong><br />
Russia (based on <strong>in</strong>formation from 2005), the<br />
majority occurred <strong>in</strong> the Komi Republic accord<strong>in</strong>g<br />
to a pre-2000 estimate [1] . Although undoubtedly<br />
important <strong>in</strong> terms of size, the Russian population<br />
Table 1.<br />
Summary of Global<br />
and European Red<br />
List assessments and<br />
threats listed for the<br />
Wolver<strong>in</strong>e.<br />
Table 2.<br />
Latest approximate<br />
population<br />
estimates for the<br />
Wolver<strong>in</strong>e <strong>in</strong> Europe<br />
and for European<br />
populations.<br />
Please note that<br />
the estimate for<br />
European Russia may<br />
be outdated.<br />
Estimate Year assessed Reference<br />
Global No data -<br />
[2]<br />
Europe 2,630–2,640 2005–11<br />
[11, 18]<br />
% of global population No data<br />
European Russia 1,400 2005<br />
[11]<br />
F<strong>in</strong>land 165–175 2011<br />
[18]<br />
Norway 385 2011<br />
[18]<br />
Sweden 680 2011<br />
[18]<br />
121