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130925-studie-wildlife-comeback-in-europe

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Estimate Year assessed Reference<br />

Global No data -<br />

-<br />

Europe 2,443,035 2002–2010<br />

[9]<br />

% of global population No data<br />

Austria 140,000–190,000 2005<br />

[40]<br />

Belgium (Wallonia) 10,000 2006<br />

[41]<br />

Belarus 4,890 2003<br />

[1]<br />

Bosnia and Herzegov<strong>in</strong>a No data - -<br />

Bulgaria 16,264 2002<br />

[1]<br />

Croatia 9,600 2002<br />

[23]<br />

Czech Republic 25,000 2004<br />

[14]<br />

Denmark 12,000–14,000 2002<br />

[42]<br />

Estonia 1,550 2005<br />

[15]<br />

France 35,000–45,000 2000<br />

[11]<br />

Germany 150,000–180,000 2006<br />

[43]<br />

Greece 130 2001<br />

[1]<br />

Hungary 74,130 2005<br />

[44]<br />

Ireland 3,000–4,000 2010<br />

[45]<br />

Italy 62,913 2005<br />

[20]<br />

Latvia 28,400 2005<br />

[15]<br />

Lithuania 12,600 2005<br />

[15]<br />

Luxembourg 3,192 2003<br />

[1]<br />

Macedonia 300 2002<br />

[1]<br />

Moldova 429 2003<br />

[1]<br />

Montenegro No data - -<br />

Netherlands 2,735 2010<br />

[46]<br />

Norway 130,000 2004<br />

[12]<br />

Poland 141,000 2005<br />

[22]<br />

Portugal 15,000–20,000 2010<br />

[19]<br />

Romania 36,100 2006<br />

[47]<br />

Russia (European) 17,630 2006<br />

[1]<br />

Serbia 5,000 2007<br />

[48]<br />

Slovakia 38,000 2004<br />

[48]<br />

Slovenia 10,000–14,000 2010<br />

[17]<br />

Spa<strong>in</strong> >800,000 2010<br />

[49]<br />

Sweden >10,000 2010<br />

[13]<br />

Switzerland 25,647 2004<br />

[16]<br />

Ukra<strong>in</strong>e 14,431 2006<br />

[1]<br />

United K<strong>in</strong>gdom ~400,000 2007<br />

[50]<br />

Table 2.<br />

Latest population<br />

estimates for the<br />

Red deer globally,<br />

<strong>in</strong> Europe and<br />

for European<br />

populations.<br />

libria <strong>in</strong> the sex ratio and, ultimately, changes <strong>in</strong><br />

the genetic structure through selective cull<strong>in</strong>g<br />

(male-biased hunt<strong>in</strong>g or cull<strong>in</strong>g of undesirable<br />

phenotypes) [34–36] . Grow<strong>in</strong>g habitat fragmentation<br />

caused by urban expansion and agricultural <strong>in</strong>tensification<br />

also represents a problem <strong>in</strong> parts of the<br />

species’ European range at present [7] .<br />

Abundance and distribution:<br />

current status<br />

While no data are available for global population<br />

size, the IUCN estimates a total of almost 2.5 million<br />

<strong>in</strong>dividuals <strong>in</strong> Europe, with the majority of these<br />

occurr<strong>in</strong>g <strong>in</strong> Spa<strong>in</strong> (32%), the UK (16%), Germany,<br />

Austria and Poland (6% each) and Norway (5%).<br />

At present, the Red deer is widely distributed<br />

throughout most of the European cont<strong>in</strong>ent, with<br />

the exception of northern Scand<strong>in</strong>avia, F<strong>in</strong>land<br />

and Iceland [2] . However, its range is likely to be<br />

patchier than <strong>in</strong>dicated on available maps [1] . For<br />

example, <strong>in</strong> Germany, the species is restricted to<br />

specific areas and rarely occurs outside of these [43] .<br />

Additionally, there is <strong>in</strong>creas<strong>in</strong>g fragmentation<br />

of populations <strong>in</strong> central Europe [1] . Traditionally<br />

considered a woodland species, Red deer has<br />

expanded <strong>in</strong>to a variety of habitats, <strong>in</strong>clud<strong>in</strong>g open<br />

moorland <strong>in</strong> the UK [45] , with densities vary<strong>in</strong>g with<br />

habitat quality [51] .<br />

Based on population numbers, Red deer is the<br />

third most common ungulate <strong>in</strong> Europe, with<br />

particularly high numbers <strong>in</strong> Spa<strong>in</strong> and the UK [9]<br />

(Table 2). However, <strong>in</strong> some parts of Europe, the<br />

species has not yet recovered from past population<br />

decl<strong>in</strong>es. For example, it is listed as Endangered <strong>in</strong><br />

the Red Data Book of Threatened Vertebrates of<br />

Greece [38] , Vulnerable <strong>in</strong> Serbia [39] and Sweden [52] ,<br />

and the subspecies corsicanus is considered Endangered<br />

on the French National Red List [53] . The<br />

species is thought to be Ext<strong>in</strong>ct <strong>in</strong> Albania [26] , and<br />

estimated at only a few hundred <strong>in</strong>dividuals <strong>in</strong><br />

Macedonia [1] .<br />

Abundance and distribution: changes<br />

Accord<strong>in</strong>g to available range <strong>in</strong>formation, the<br />

Red deer has expanded its area of occurrence<br />

by 190% s<strong>in</strong>ce 1955 , spread<strong>in</strong>g <strong>in</strong>to surround<strong>in</strong>g<br />

areas from refugia and now occupy<strong>in</strong>g nearly half<br />

of its historical range <strong>in</strong> the 1800s (Figure 1a). For<br />

example, the British Isles have seen range <strong>in</strong>creases<br />

of between 0.3% (United K<strong>in</strong>gdom [10] ) and 7% per<br />

year (Ireland [54] ). However, the expansion depicted<br />

for the UK <strong>in</strong> Figure 1b is likely an exaggeration [55]<br />

result<strong>in</strong>g from differences <strong>in</strong> the resolution of the<br />

two maps. Similarly, the range depicted for the<br />

1950s <strong>in</strong> Spa<strong>in</strong> is larger than the actual distribution<br />

at this po<strong>in</strong>t <strong>in</strong> time, as the species was reduced<br />

to small patches <strong>in</strong> Extremadura, Sierra Morena<br />

and Montes de Toledo [31] . The majority of extant<br />

populations <strong>in</strong> the north of Spa<strong>in</strong>, Portugal and<br />

some southernmost ranges have resulted from<br />

translocations from these rema<strong>in</strong><strong>in</strong>g stocks [31] . In<br />

addition, current range may be overestimated <strong>in</strong><br />

some countries, for example <strong>in</strong> Germany, where<br />

the species is legally conf<strong>in</strong>ed to Red deer areas<br />

(“Rotwildgebiete”), outside of which <strong>in</strong>dividuals<br />

are generally shot [43] . Despite the overall positive<br />

developments, there has also been some<br />

contraction at a sub-regional level or national level,<br />

most notably <strong>in</strong> the already threatened population<br />

<strong>in</strong> Greece (Figure 1a).<br />

The expansion <strong>in</strong> range is also reflected <strong>in</strong> the<br />

change <strong>in</strong> population size over a similar period.<br />

Overall, the species has experienced an <strong>in</strong>crease<br />

76

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