130925-studie-wildlife-comeback-in-europe
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Estimate Year assessed Reference<br />
Global >1,036,226 2012<br />
[11]<br />
Europe (m<strong>in</strong>imum) >337,539 2003–12<br />
[6, 12–18]<br />
% of global population 33%<br />
Austria >3,000 2008<br />
[12]<br />
Belarus 24,000 2003<br />
[6]<br />
Belgium 200–250 2003<br />
[6]<br />
Bosnia and Herzegov<strong>in</strong>a Unknown - -<br />
Croatia 180 2003<br />
[6]<br />
Czech Republic 500 2003<br />
[6]<br />
Denmark 188 2011<br />
[13]<br />
Estonia 11,000 2003<br />
[6]<br />
F<strong>in</strong>land 2,000 2003<br />
[6]<br />
France 14,000 2011<br />
[14]<br />
Germany >25000 2013<br />
[12]<br />
Hungary 500 2007<br />
[15]<br />
Latvia >100,000 2003<br />
[6]<br />
Lithuania 50,000–70,000 2003<br />
[6]<br />
Netherlands 507 2010<br />
[16]<br />
Norway 70,000 2003<br />
[6]<br />
Poland 18,000–23,000 2003<br />
[6]<br />
Romania >170 2003<br />
[6]<br />
Russia (European) Unknown - -<br />
Serbia 30 2003<br />
[6]<br />
Slovakia >500 2003<br />
[6]<br />
Spa<strong>in</strong> 18 2003<br />
[6]<br />
Sweden >100,000 2003<br />
[6]<br />
Switzerland 1,600 2007/8<br />
[17]<br />
Ukra<strong>in</strong>e 6,000 2003<br />
[6]<br />
United K<strong>in</strong>gdom 146 2012<br />
[18]<br />
1845 (which was only effective from 1899), numbers<br />
recovered to 7,000 <strong>in</strong> 1919; however, subsequent<br />
over-hunt<strong>in</strong>g caused population decl<strong>in</strong>es despite<br />
re<strong>in</strong>troductions <strong>in</strong> the 1920s [5] . In the early 1940s,<br />
<strong>in</strong>dividuals immigrated to eastern Norway from<br />
the re<strong>in</strong>troduced Swedish population along the<br />
Trysil watershed [5] . This natural spread cont<strong>in</strong>ued,<br />
becom<strong>in</strong>g a significant factor <strong>in</strong> the 1970s, and<br />
this, <strong>in</strong> comb<strong>in</strong>ation with re<strong>in</strong>troductions <strong>in</strong> the<br />
1950s and 1960s <strong>in</strong> Norway, led to considerable<br />
<strong>in</strong>creases <strong>in</strong> number and range, leav<strong>in</strong>g the species<br />
to occupy the country <strong>in</strong> two disjunct populations<br />
(Figure 1a) [5] .<br />
Hunted to ext<strong>in</strong>ction <strong>in</strong> 1938, Lithuania provides<br />
ideal habitat for the Eurasian beaver [5] . Re<strong>in</strong>troductions<br />
between 1947 and 1959, coupled with<br />
immigration from Latvia, Belarus and Kal<strong>in</strong><strong>in</strong>grad<br />
<strong>in</strong> Russia, has led to a constant <strong>in</strong>crease s<strong>in</strong>ce the<br />
end of the 1940s [5] . Even though range expansion<br />
halted <strong>in</strong> the mid-1970s, when the beaver occupied<br />
the entire country [5] , <strong>in</strong>creases have cont<strong>in</strong>ued <strong>in</strong><br />
the late 1990s due to the post-Soviet abandonment<br />
of dra<strong>in</strong>ed farmland lead<strong>in</strong>g to scrub regrowth,<br />
which comb<strong>in</strong>ed with exist<strong>in</strong>g dra<strong>in</strong>age ditches<br />
provide ideal beaver habitat [2] .<br />
Abundance and distribution: changes<br />
Table 2.<br />
Latest population<br />
estimates for the<br />
Eurasian beaver<br />
globally, <strong>in</strong> Europe<br />
and for European<br />
populations. Please<br />
note that there<br />
was no <strong>in</strong>formation<br />
available for Bosnia<br />
and Herzegov<strong>in</strong>a, and<br />
European Russia.<br />
is currently found throughout Latvia except the<br />
northwestern part [5] .<br />
The beaver occurs <strong>in</strong> all of Sweden except the<br />
south, northwest, and the area around Uppsala<br />
north of Stockholm, and this population forms<br />
part of the cont<strong>in</strong>uous distribution of the species<br />
from the Baltic to the Atlantic [5] . The population,<br />
which was extirpated <strong>in</strong> 1871 [6] , was re-established<br />
entirely through re<strong>in</strong>troductions [2] . Follow<strong>in</strong>g<br />
re<strong>in</strong>troductions of Norwegian beavers from<br />
1922 at 19 sites, the species was able to expand<br />
reasonably quickly due to limited barriers<br />
result<strong>in</strong>g from the topological characteristics of<br />
the country [5] . After rapid range extension, the<br />
characteristic population explosion took place <strong>in</strong><br />
the 1970s, result<strong>in</strong>g <strong>in</strong> an estimated population<br />
of over 100,000 <strong>in</strong>dividuals spread across what<br />
is now a cont<strong>in</strong>uous range [5] . Populations are<br />
completely protected <strong>in</strong> the northernmost<br />
prov<strong>in</strong>ce Norrbotten, <strong>in</strong> the eastern Uppsala and<br />
Stockholm prov<strong>in</strong>ces, and <strong>in</strong> the south, while<br />
seasonal unrestricted hunt<strong>in</strong>g elsewhere results <strong>in</strong><br />
an annual harvest of around 6% per annum [5] .<br />
Grow<strong>in</strong>g from a remnant population of around<br />
100 <strong>in</strong>dividuals [5] , the Norwegian beaver has<br />
provided source stock for all Scand<strong>in</strong>avian populations<br />
and now represents up to 21% of the species<br />
<strong>in</strong> Europe (Table 2). After receiv<strong>in</strong>g protection <strong>in</strong><br />
In 1955, the Eurasian beaver occurred <strong>in</strong> over 25<br />
dist<strong>in</strong>ct populations <strong>in</strong> eastern Germany, the south<br />
of France, eastern Europe and Scand<strong>in</strong>avia (Figure 1a),<br />
occupy<strong>in</strong>g an area of over 256,000 km 2 . By 2013, the<br />
range had <strong>in</strong>creased by 650% to nearly 1,670,000<br />
km 2 , add<strong>in</strong>g territory across the European cont<strong>in</strong>ent<br />
(Figures 1a and b). While the species seem<strong>in</strong>gly lost<br />
ground <strong>in</strong> the south of France, eastern Germany,<br />
Norway, northern Sweden and Ukra<strong>in</strong>e, this is likely<br />
to be an artefact of the difference <strong>in</strong> map resolution.<br />
The 1955 map is very simplified, giv<strong>in</strong>g the impression<br />
that the beaver has contracted compared to the more<br />
detailed 2013 range. However, it is possible that even<br />
the present day map, although broadly accurate,<br />
represents an underestimate of distribution due<br />
to the rapidly chang<strong>in</strong>g situation of the species<br />
across the cont<strong>in</strong>ent [1] , which would lead to greater<br />
expansion s<strong>in</strong>ce 1955 than depicted <strong>in</strong> Figure 1b.<br />
Range loss may, however, be a reality <strong>in</strong> F<strong>in</strong>land,<br />
where the beaver is experienc<strong>in</strong>g <strong>in</strong>terspecific<br />
competition with the <strong>in</strong>troduced North American<br />
beaver (Castor canadensis) [1] . Despite this, previously<br />
disjunct populations have been connected, creat<strong>in</strong>g<br />
large ranges <strong>in</strong> central and eastern Europe, and <strong>in</strong><br />
Norway and Sweden, and creat<strong>in</strong>g new populations<br />
<strong>in</strong> countries where the species had been absent for<br />
many centuries, for example Spa<strong>in</strong> and the United<br />
K<strong>in</strong>gdom (Figures 1a and b).<br />
152