130925-studie-wildlife-comeback-in-europe
Create successful ePaper yourself
Turn your PDF publications into a flip-book with our unique Google optimized e-Paper software.
Habitat preferences and general densities<br />
The Northern chamois occurs <strong>in</strong> a number of<br />
habitats <strong>in</strong>clud<strong>in</strong>g alp<strong>in</strong>e meadows, open rocky<br />
areas, mixed broadleaf woodland and coniferous<br />
woodland <strong>in</strong> steep, rocky, mounta<strong>in</strong>ous areas [5] .<br />
Dur<strong>in</strong>g w<strong>in</strong>ter, it moves from alp<strong>in</strong>e meadows<br />
to lower-ly<strong>in</strong>g, forested areas [6] . When alarmed,<br />
chamois retreat to highly <strong>in</strong>accessible places,<br />
mak<strong>in</strong>g leaps of 2 m <strong>in</strong> height and 6 m <strong>in</strong> length [3] .<br />
Legal protection and conservation status<br />
In Europe, the subspecies R. r. balcanica and tatrica<br />
are listed on Annexes II and IV of the EU Habitats<br />
Directive, while the whole species is <strong>in</strong>cluded on<br />
Annex V of the Directive [7] and Appendix III of the<br />
Bern Convention [8] . Conservation actions for all<br />
subspecies <strong>in</strong>clude ensur<strong>in</strong>g susta<strong>in</strong>able harvest,<br />
reduc<strong>in</strong>g poach<strong>in</strong>g, reduc<strong>in</strong>g human disturbance,<br />
protect<strong>in</strong>g the genetic <strong>in</strong>tegrity of populations,<br />
and monitor<strong>in</strong>g, especially of vulnerable populations<br />
[9] . There have also been re<strong>in</strong>troductions<br />
of R. r. cartusiana [9] and <strong>in</strong> some areas protected<br />
areas have been set up. The hunt<strong>in</strong>g of the species<br />
is widespread and common, and cull<strong>in</strong>g is used<br />
to control population numbers, e.g. <strong>in</strong> France [4] .<br />
Both at a global and European level, the Northern<br />
chamois is classified as Least Concern with an<br />
unknown population trend (Table 2). However,<br />
a number of the subspecies are believed to be<br />
decreas<strong>in</strong>g and are listed as Critically Endangered<br />
or Vulnerable due to various threats (Table 2).<br />
Abundance and distribution:<br />
current status<br />
In terms of population size, an estimate from<br />
2004/5 puts the European population at 485,580<br />
<strong>in</strong>dividuals, but as there are no reliable global<br />
estimates available, it is not possible to ascerta<strong>in</strong><br />
the proportion accounted for by this population<br />
(Table 3). With<strong>in</strong> Europe, the largest populations<br />
occur <strong>in</strong> the Alp<strong>in</strong>e arc countries Austria (31%),<br />
Italy (28%), Switzerland (19%) and France (13%),<br />
and mostly as the subspecies R. r. rupicapra. The<br />
majority of extant populations <strong>in</strong> central Europe<br />
are re<strong>in</strong>troduced [6] .<br />
R. r. rupicapra represents the most numerous<br />
of the subspecies, occurr<strong>in</strong>g throughout the<br />
Austrian, Italian, Swiss and French Alps [6] . In<br />
Austria, the species is <strong>in</strong>creas<strong>in</strong>gly found <strong>in</strong><br />
suboptimal habitat [6] , which suggests that it is<br />
do<strong>in</strong>g well here. Measures are <strong>in</strong> place to manage<br />
sarcoptic mange [6] , and there are abundance-dependent<br />
hunt<strong>in</strong>g quotas [6] . The chamois is also<br />
hunted <strong>in</strong> Italy [6] . While present <strong>in</strong> almost all<br />
of Switzerland historically [13] , the species now<br />
Subspecies Distribution Population size and trend<br />
balcanica<br />
[9]<br />
Mounta<strong>in</strong> regions of Albania<br />
Bulgaria’s four ma<strong>in</strong> massifs<br />
Thousands of <strong>in</strong>dividuals<br />
Decl<strong>in</strong><strong>in</strong>g <strong>in</strong> all subpopulations<br />
6 populations on 11 mounta<strong>in</strong>s<br />
<strong>in</strong> Greece (Mount Rodopi <strong>in</strong><br />
the northeast and the Epirus<br />
mounta<strong>in</strong>s <strong>in</strong> the northwest to<br />
Mount Giona <strong>in</strong> central Greece)<br />
carpatica<br />
[1, 9]<br />
Transylvanian Alps<br />
Carpathians<br />
9,000 <strong>in</strong>dividuals <strong>in</strong> 1990<br />
Increas<strong>in</strong>g <strong>in</strong> Romania<br />
successful re<strong>in</strong>troductions<br />
cartusiana<br />
[1]<br />
Chartreuse limestone massif<br />
around Grenoble <strong>in</strong> France<br />
Western edge of French Alps<br />
300–400 <strong>in</strong>dividuals <strong>in</strong> 1970s<br />
150 <strong>in</strong>dividuals <strong>in</strong> 1986–7<br />
Recent estimate: 2,000 <strong>in</strong>dividuals<br />
rupicapra<br />
[1, 10]<br />
Alps (Austria, Germany, eastern<br />
France)<br />
Comprises majority of global population<br />
Widespread and abundant<br />
Cull<strong>in</strong>g <strong>in</strong> Swiss Alps and Jura rose from<br />
4,000 <strong>in</strong>dividuals (1950) to 17,000 (2000)<br />
tatrica<br />
[1, 9, 11]<br />
Tatra mounta<strong>in</strong>s of Poland and<br />
Slovakia<br />
Introduced <strong>in</strong>to the low Tatras<br />
<strong>in</strong> Slovakia<br />
only occurs <strong>in</strong> the Alps and parts of the Jura<br />
mounta<strong>in</strong>s [6] , where it was re<strong>in</strong>troduced between<br />
1950 and 1962 [14] . The first federal hunt<strong>in</strong>g law<br />
was established <strong>in</strong> 1875 and as a result, populations<br />
recovered swiftly [15] .<br />
In France, R. r. rupicapra occurs <strong>in</strong> the Alp<strong>in</strong>e<br />
region, the Jura and Vosges mounta<strong>in</strong>s and the<br />
Massif Central, but the country is also home<br />
to R. r. cartusiana <strong>in</strong> the Chartreuse limestone<br />
massif [6] . Hunt<strong>in</strong>g quotas have been <strong>in</strong> place s<strong>in</strong>ce<br />
1990 [4] , with an <strong>in</strong>itial reduction <strong>in</strong> cull<strong>in</strong>g lead<strong>in</strong>g<br />
to a recovery, which was followed by a subsequent<br />
higher cull<strong>in</strong>g quota [4] . The cartusiana subspecies<br />
recovered from food competition with other<br />
species from 250 <strong>in</strong>dividuals <strong>in</strong> 1972 to over 770 by<br />
1997 [16] . However, food competition is still a major<br />
threat [16] , as is hybridisation with the spread<strong>in</strong>g<br />
Alp<strong>in</strong>e chamois, which was <strong>in</strong>troduced to the<br />
northern end of the massif [16] .<br />
R. r. tatrica is present <strong>in</strong> two locations: one <strong>in</strong><br />
the High Tatra mounta<strong>in</strong>s <strong>in</strong> northern Slovakia<br />
and southern Poland, and a second, re<strong>in</strong>troduced<br />
population <strong>in</strong> the Low Tatra <strong>in</strong> Slovakia [11] .<br />
Decl<strong>in</strong>es after the first and second world wars to<br />
300 and 132–230 <strong>in</strong>dividuals [11] were followed by<br />
<strong>in</strong>itial recovery and then further decl<strong>in</strong>es, which<br />
cont<strong>in</strong>ued steadily from the 1960s until today [11] .<br />
They are attributed to altered age structure and<br />
sex ratio <strong>in</strong>duced by selective shoot<strong>in</strong>g, severe<br />
weather conditions, human disturbance through<br />
tourism and air traffic, predation and parasitism [11] .<br />
Censuses <strong>in</strong> 1999 counted 220 and 120–130<br />
<strong>in</strong>dividuals <strong>in</strong> the two populations respectively [11] .<br />
Poach<strong>in</strong>g and potential hybridisation with <strong>in</strong>troduced<br />
Alp<strong>in</strong>e chamois have been highlighted as<br />
the most urgent threats to address [11] .<br />
Decl<strong>in</strong><strong>in</strong>g steadily s<strong>in</strong>ce 1960<br />
220 <strong>in</strong>dividuals <strong>in</strong> 1999<br />