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Habitat preferences and general densities<br />

The Northern chamois occurs <strong>in</strong> a number of<br />

habitats <strong>in</strong>clud<strong>in</strong>g alp<strong>in</strong>e meadows, open rocky<br />

areas, mixed broadleaf woodland and coniferous<br />

woodland <strong>in</strong> steep, rocky, mounta<strong>in</strong>ous areas [5] .<br />

Dur<strong>in</strong>g w<strong>in</strong>ter, it moves from alp<strong>in</strong>e meadows<br />

to lower-ly<strong>in</strong>g, forested areas [6] . When alarmed,<br />

chamois retreat to highly <strong>in</strong>accessible places,<br />

mak<strong>in</strong>g leaps of 2 m <strong>in</strong> height and 6 m <strong>in</strong> length [3] .<br />

Legal protection and conservation status<br />

In Europe, the subspecies R. r. balcanica and tatrica<br />

are listed on Annexes II and IV of the EU Habitats<br />

Directive, while the whole species is <strong>in</strong>cluded on<br />

Annex V of the Directive [7] and Appendix III of the<br />

Bern Convention [8] . Conservation actions for all<br />

subspecies <strong>in</strong>clude ensur<strong>in</strong>g susta<strong>in</strong>able harvest,<br />

reduc<strong>in</strong>g poach<strong>in</strong>g, reduc<strong>in</strong>g human disturbance,<br />

protect<strong>in</strong>g the genetic <strong>in</strong>tegrity of populations,<br />

and monitor<strong>in</strong>g, especially of vulnerable populations<br />

[9] . There have also been re<strong>in</strong>troductions<br />

of R. r. cartusiana [9] and <strong>in</strong> some areas protected<br />

areas have been set up. The hunt<strong>in</strong>g of the species<br />

is widespread and common, and cull<strong>in</strong>g is used<br />

to control population numbers, e.g. <strong>in</strong> France [4] .<br />

Both at a global and European level, the Northern<br />

chamois is classified as Least Concern with an<br />

unknown population trend (Table 2). However,<br />

a number of the subspecies are believed to be<br />

decreas<strong>in</strong>g and are listed as Critically Endangered<br />

or Vulnerable due to various threats (Table 2).<br />

Abundance and distribution:<br />

current status<br />

In terms of population size, an estimate from<br />

2004/5 puts the European population at 485,580<br />

<strong>in</strong>dividuals, but as there are no reliable global<br />

estimates available, it is not possible to ascerta<strong>in</strong><br />

the proportion accounted for by this population<br />

(Table 3). With<strong>in</strong> Europe, the largest populations<br />

occur <strong>in</strong> the Alp<strong>in</strong>e arc countries Austria (31%),<br />

Italy (28%), Switzerland (19%) and France (13%),<br />

and mostly as the subspecies R. r. rupicapra. The<br />

majority of extant populations <strong>in</strong> central Europe<br />

are re<strong>in</strong>troduced [6] .<br />

R. r. rupicapra represents the most numerous<br />

of the subspecies, occurr<strong>in</strong>g throughout the<br />

Austrian, Italian, Swiss and French Alps [6] . In<br />

Austria, the species is <strong>in</strong>creas<strong>in</strong>gly found <strong>in</strong><br />

suboptimal habitat [6] , which suggests that it is<br />

do<strong>in</strong>g well here. Measures are <strong>in</strong> place to manage<br />

sarcoptic mange [6] , and there are abundance-dependent<br />

hunt<strong>in</strong>g quotas [6] . The chamois is also<br />

hunted <strong>in</strong> Italy [6] . While present <strong>in</strong> almost all<br />

of Switzerland historically [13] , the species now<br />

Subspecies Distribution Population size and trend<br />

balcanica<br />

[9]<br />

Mounta<strong>in</strong> regions of Albania<br />

Bulgaria’s four ma<strong>in</strong> massifs<br />

Thousands of <strong>in</strong>dividuals<br />

Decl<strong>in</strong><strong>in</strong>g <strong>in</strong> all subpopulations<br />

6 populations on 11 mounta<strong>in</strong>s<br />

<strong>in</strong> Greece (Mount Rodopi <strong>in</strong><br />

the northeast and the Epirus<br />

mounta<strong>in</strong>s <strong>in</strong> the northwest to<br />

Mount Giona <strong>in</strong> central Greece)<br />

carpatica<br />

[1, 9]<br />

Transylvanian Alps<br />

Carpathians<br />

9,000 <strong>in</strong>dividuals <strong>in</strong> 1990<br />

Increas<strong>in</strong>g <strong>in</strong> Romania<br />

successful re<strong>in</strong>troductions<br />

cartusiana<br />

[1]<br />

Chartreuse limestone massif<br />

around Grenoble <strong>in</strong> France<br />

Western edge of French Alps<br />

300–400 <strong>in</strong>dividuals <strong>in</strong> 1970s<br />

150 <strong>in</strong>dividuals <strong>in</strong> 1986–7<br />

Recent estimate: 2,000 <strong>in</strong>dividuals<br />

rupicapra<br />

[1, 10]<br />

Alps (Austria, Germany, eastern<br />

France)<br />

Comprises majority of global population<br />

Widespread and abundant<br />

Cull<strong>in</strong>g <strong>in</strong> Swiss Alps and Jura rose from<br />

4,000 <strong>in</strong>dividuals (1950) to 17,000 (2000)<br />

tatrica<br />

[1, 9, 11]<br />

Tatra mounta<strong>in</strong>s of Poland and<br />

Slovakia<br />

Introduced <strong>in</strong>to the low Tatras<br />

<strong>in</strong> Slovakia<br />

only occurs <strong>in</strong> the Alps and parts of the Jura<br />

mounta<strong>in</strong>s [6] , where it was re<strong>in</strong>troduced between<br />

1950 and 1962 [14] . The first federal hunt<strong>in</strong>g law<br />

was established <strong>in</strong> 1875 and as a result, populations<br />

recovered swiftly [15] .<br />

In France, R. r. rupicapra occurs <strong>in</strong> the Alp<strong>in</strong>e<br />

region, the Jura and Vosges mounta<strong>in</strong>s and the<br />

Massif Central, but the country is also home<br />

to R. r. cartusiana <strong>in</strong> the Chartreuse limestone<br />

massif [6] . Hunt<strong>in</strong>g quotas have been <strong>in</strong> place s<strong>in</strong>ce<br />

1990 [4] , with an <strong>in</strong>itial reduction <strong>in</strong> cull<strong>in</strong>g lead<strong>in</strong>g<br />

to a recovery, which was followed by a subsequent<br />

higher cull<strong>in</strong>g quota [4] . The cartusiana subspecies<br />

recovered from food competition with other<br />

species from 250 <strong>in</strong>dividuals <strong>in</strong> 1972 to over 770 by<br />

1997 [16] . However, food competition is still a major<br />

threat [16] , as is hybridisation with the spread<strong>in</strong>g<br />

Alp<strong>in</strong>e chamois, which was <strong>in</strong>troduced to the<br />

northern end of the massif [16] .<br />

R. r. tatrica is present <strong>in</strong> two locations: one <strong>in</strong><br />

the High Tatra mounta<strong>in</strong>s <strong>in</strong> northern Slovakia<br />

and southern Poland, and a second, re<strong>in</strong>troduced<br />

population <strong>in</strong> the Low Tatra <strong>in</strong> Slovakia [11] .<br />

Decl<strong>in</strong>es after the first and second world wars to<br />

300 and 132–230 <strong>in</strong>dividuals [11] were followed by<br />

<strong>in</strong>itial recovery and then further decl<strong>in</strong>es, which<br />

cont<strong>in</strong>ued steadily from the 1960s until today [11] .<br />

They are attributed to altered age structure and<br />

sex ratio <strong>in</strong>duced by selective shoot<strong>in</strong>g, severe<br />

weather conditions, human disturbance through<br />

tourism and air traffic, predation and parasitism [11] .<br />

Censuses <strong>in</strong> 1999 counted 220 and 120–130<br />

<strong>in</strong>dividuals <strong>in</strong> the two populations respectively [11] .<br />

Poach<strong>in</strong>g and potential hybridisation with <strong>in</strong>troduced<br />

Alp<strong>in</strong>e chamois have been highlighted as<br />

the most urgent threats to address [11] .<br />

Decl<strong>in</strong><strong>in</strong>g steadily s<strong>in</strong>ce 1960<br />

220 <strong>in</strong>dividuals <strong>in</strong> 1999<br />

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