130925-studie-wildlife-comeback-in-europe
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3.8. Red deer<br />
Cervus elaphus<br />
Summary<br />
The Red deer has a wide distribution across most<br />
of the European cont<strong>in</strong>ent, with the exception of<br />
northern Scand<strong>in</strong>avia, F<strong>in</strong>land and Iceland, where<br />
it is limited by severe climate. Decl<strong>in</strong>es and local<br />
ext<strong>in</strong>ctions occurred throughout the species’<br />
native range from the 16 th century due to overexploitation,<br />
habitat loss and competition with<br />
livestock. Through changes <strong>in</strong> legislation, hunt<strong>in</strong>g,<br />
habitat, and predator and competitor levels, as<br />
well as translocations, re<strong>in</strong>troductions and recolonisation,<br />
the cervid has grown <strong>in</strong> abundance and<br />
range, and now represents the greatest biomass of<br />
any ungulate <strong>in</strong> Europe.<br />
Background<br />
General description of the species<br />
The Red deer (Cervus elaphus) is the most widely<br />
distributed deer species <strong>in</strong> the world, with a large<br />
but patchy distribution across Eurasia and also<br />
reach<strong>in</strong>g <strong>in</strong>to northern Africa [1, 2] , be<strong>in</strong>g limited<br />
only by severe w<strong>in</strong>ter climate [3] . It also occurs <strong>in</strong><br />
<strong>in</strong>troduced populations <strong>in</strong> parts of the southern<br />
hemisphere such as Chile, Argent<strong>in</strong>a, Australia<br />
and New Zealand [4] . Once considered a s<strong>in</strong>gle<br />
species, Red deer is now divided <strong>in</strong>to six to ten<br />
subspecies globally [4] . It is the fourth largest<br />
ungulate [5] and second largest deer on the European<br />
cont<strong>in</strong>ent [6] . Although active throughout day and<br />
night, peaks of activity occur at twilight due to<br />
human activity [6] . As an <strong>in</strong>termediate feeder with<br />
a large rumen, the Red deer consumes a variety<br />
of plants <strong>in</strong>clud<strong>in</strong>g grasses, sedges, browse, fruits<br />
and seeds; the exact composition differs between<br />
males and females, and habitats [4] . Females and<br />
young live <strong>in</strong> small matril<strong>in</strong>eal herds but gather<br />
<strong>in</strong>to larger groups <strong>in</strong> w<strong>in</strong>ter, while stags live a<br />
solitary life, except <strong>in</strong> summer when they may<br />
occur <strong>in</strong> all-male herds, and dur<strong>in</strong>g the rut <strong>in</strong> late<br />
summer when they gather harems [4] . One calf<br />
is usually born <strong>in</strong> May or June after a gestation<br />
period of around 8.5 months [4] . Young reach sexual<br />
maturity between 1.5 and 2.5 years depend<strong>in</strong>g on<br />
habitat quality [4] . The natural maximum lifespan<br />
of the species is 17–18 years, although <strong>in</strong>dividuals<br />
as old as 26 years have been recorded [4] .<br />
Distribution <strong>in</strong> Europe<br />
The species appeared <strong>in</strong> Europe <strong>in</strong> the late Early<br />
Pleistocene around 900,000 years ago [4] and was<br />
able to persist <strong>in</strong> southern Europe (Iberia, southwestern<br />
France, Italy, Balkans, Greece) and east<br />
of the Carpathians <strong>in</strong> Moldavia dur<strong>in</strong>g the Last<br />
Glacial Maximum [7, 8] . The clades present today can<br />
be traced back to different refugia: the western and<br />
eastern European l<strong>in</strong>eages (scoticus, atlanticus,<br />
elaphus, hippelaphus and hispanicus) stem from<br />
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