130925-studie-wildlife-comeback-in-europe
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3.3. Iberian ibex<br />
Capra pyrenaica<br />
Summary<br />
The Iberian ibex decl<strong>in</strong>ed due to over-exploitation,<br />
poach<strong>in</strong>g, <strong>in</strong>fection and <strong>in</strong>ter-species competition.<br />
In contrast, legal protection, translocations and<br />
re<strong>in</strong>troductions, and new habitat result<strong>in</strong>g from<br />
rural abandonment have had positive effects on<br />
the species’ distribution and abundance. However,<br />
there are differences <strong>in</strong> ecology between the two<br />
rema<strong>in</strong><strong>in</strong>g subspecies, and future management<br />
strategies will have to be devised accord<strong>in</strong>gly.<br />
Background<br />
General description of the species<br />
The Iberian ibex (Capra pyrenaica), a familiar and<br />
popular species due to its occurrence <strong>in</strong> close<br />
proximity to humans, is endemic to the Iberian<br />
pen<strong>in</strong>sula [1] . As a mixed feeder, it browses or grazes<br />
depend<strong>in</strong>g on the availability of plants, and diet is<br />
<strong>in</strong>fluenced by altitude [2] , geographic location [3] and<br />
season [4] . Based on the small genetic distance between<br />
Iberian and Alp<strong>in</strong>e ibex (Capra ibex) [5] as well as paleontological<br />
descriptions found <strong>in</strong> Germany [6] , the species<br />
are believed to have split follow<strong>in</strong>g a wave of Capra<br />
immigration <strong>in</strong>to Europe 300,000 years ago [5] .<br />
Distribution <strong>in</strong> Europe<br />
Once widely distributed throughout southwest<br />
France, Spa<strong>in</strong>, Andorra, and Portugal, the ibex<br />
decreased significantly over the past 200 years<br />
due to over-exploitation and habitat loss [7–9] . It<br />
formerly existed <strong>in</strong> four subspecies [1] . C. p. lusitanica,<br />
which <strong>in</strong>habited northwest Portugal and Galicia<br />
<strong>in</strong> Spa<strong>in</strong>, died out at the end of the 19 th century [10] ;<br />
and C. p. pyrenaica, which was abundant <strong>in</strong> the<br />
Pyrenees <strong>in</strong> the Middle Ages [11] and rema<strong>in</strong>ed <strong>in</strong><br />
the Spanish territory until its ext<strong>in</strong>ction <strong>in</strong> 2000<br />
due to overhunt<strong>in</strong>g, agricultural development and<br />
expansion, and habitat deterioration [7, 12] . Today,<br />
two subspecies rema<strong>in</strong>: C. p. hispanica <strong>in</strong> central<br />
and Mediterranean Spanish mounta<strong>in</strong> ranges, and<br />
C. p. victoriae <strong>in</strong> the northwest Iberian Pen<strong>in</strong>sula [13] .<br />
There is, however, some controversy about the<br />
subspecies classification, with some not be<strong>in</strong>g recognised<br />
[14–16] . Despite drastic reductions <strong>in</strong> the sizes of<br />
some populations [7] , ibex <strong>in</strong> Spa<strong>in</strong> have generally<br />
<strong>in</strong>creased <strong>in</strong> both number and range over the last<br />
three decades [7] . The species is now widespread <strong>in</strong> the<br />
Iberian Pen<strong>in</strong>sula [17] [7, 18]<br />
, exist<strong>in</strong>g <strong>in</strong> over 50 localities<br />
and expand<strong>in</strong>g its range <strong>in</strong>to Portugal [1, 13] .<br />
Habitat preferences and general densities<br />
The Iberian ibex prefers rocky habitats with bare,<br />
steep slopes [19] . Although it often colonises new<br />
areas rapidly through dispersal [17, 20] , the species<br />
can become displaced to less optimal habitat such<br />
as pasture-scrub land due to competition with<br />
livestock [21] .<br />
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