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from F<strong>in</strong>land to northern Ukra<strong>in</strong>e, throughout<br />

the Balkan countries, around the Carpathians <strong>in</strong><br />

eastern Europe, <strong>in</strong> central Scand<strong>in</strong>avia, along the<br />

Alps and the Italian pen<strong>in</strong>sula, and northern Spa<strong>in</strong><br />

and Portugal, with smaller populations <strong>in</strong> southcentral<br />

Spa<strong>in</strong>, and across Germany and Poland [8] .<br />

Habitat preferences and general densities<br />

Dependent on prey density and the level of human<br />

disturbance, the Grey wolf is found <strong>in</strong> a wide variety<br />

of habitats with<strong>in</strong> the Northern hemisphere [6] ,<br />

where suitable food is abundant [1] . This <strong>in</strong>cludes<br />

dense forest, open grasslands, mounta<strong>in</strong> ranges and<br />

the Arctic tundra [5] , as well as <strong>in</strong> highly human-modified<br />

habitats such as agricultural areas [9] . Wolf<br />

densities vary from 0.1 to 1 <strong>in</strong>dividual per 12 km 2 ,<br />

with the greatest numbers be<strong>in</strong>g recorded <strong>in</strong> areas<br />

where prey biomass is highest [1] .<br />

Legal protection and conservation status<br />

The Grey wolf has been protected <strong>in</strong> most of Europe<br />

s<strong>in</strong>ce 1979, when it was <strong>in</strong>cluded <strong>in</strong> Appendix II of<br />

the Bern Convention [10] . The species is now recolonis<strong>in</strong>g<br />

areas <strong>in</strong> central and western Europe such<br />

as the Western Alps [6, 11] and Scand<strong>in</strong>avia [12] . In<br />

addition, it is listed on Appendix II of CITES [13] and<br />

protected by the EC Habitat Directive [14] , with the<br />

exception of some populations <strong>in</strong> Spa<strong>in</strong>, Greece<br />

and F<strong>in</strong>land. Many countries do not enforce this<br />

level of protection and there are still significant<br />

illegal kill<strong>in</strong>gs [2] . While many wolf populations<br />

occur <strong>in</strong> protected areas [1] , these are usually too<br />

small to be of benefit to the species [12] . In many<br />

European countries compensation schemes exist<br />

for loss of livestock due to wolf predation [15] .<br />

Abundance and distribution:<br />

current status<br />

The Grey wolf’s global Red List category is Least<br />

Concern [6] due to its large range and stable global<br />

population trend [6] . However, threats to the species<br />

rema<strong>in</strong>: hunt<strong>in</strong>g and poach<strong>in</strong>g are widespread and<br />

likely the most significant threat <strong>in</strong> Europe [2] . Where<br />

human and wolf distributions overlap, livestock<br />

depredation leads to the kill<strong>in</strong>g of wolves, particularly<br />

<strong>in</strong> develop<strong>in</strong>g countries [6] . The low densities<br />

at which wolves exist make them particularly<br />

vulnerable to hunt<strong>in</strong>g as well as stochastic events [2] .<br />

In addition, habitat fragmentation is a significant<br />

threat through the isolation of small populations and<br />

the chance of dispersal to unsuitable habitats [2] . In<br />

Europe, the species is also listed as Least Concern [15] ,<br />

although the majority of regional populations are<br />

threatened due to their small size. Only the Central<br />

European and Sierra Morena populations are<br />

considered to be Critically Endangered (Table 1). In<br />

terms of population size, the current global estimate<br />

is around 200,000 <strong>in</strong>dividuals (Table 1). Europe<br />

comprises only a fraction of the species’ global distribution<br />

and accounts for a m<strong>in</strong>imum of 7% or 11,500<br />

<strong>in</strong>dividuals. Around 82% of European wolves (not<br />

<strong>in</strong>clud<strong>in</strong>g Belarus, Ukra<strong>in</strong>e and European Russia) are<br />

found <strong>in</strong> the D<strong>in</strong>aric-Balkan, Carpathian, Baltic and<br />

Iberian populations (Table 2).<br />

The D<strong>in</strong>aric-Balkan population accounts for<br />

around one-third of Europe’s wolves and covers a<br />

large area from Slovenia to northcentral Greece [15] ,<br />

<strong>in</strong>clud<strong>in</strong>g Bulgaria, Croatia, Bosnia and Herzegov<strong>in</strong>a,<br />

Serbia, Montenegro, FYR Macedonia and<br />

Albania [17] . Its conservation status is favourable due<br />

Table 1.<br />

Summary of Global<br />

and European Red<br />

List assessments and<br />

threats listed for the<br />

Grey wolf.<br />

Scale Status Population trend Justification Threats<br />

Global<br />

Least Concern Stable Wide range<br />

1. Competition with humans for livestock and wild<br />

[6]<br />

Stable population<br />

ungulates (especially <strong>in</strong> develop<strong>in</strong>g countries)<br />

2. Poison<strong>in</strong>g and persecution<br />

3. Low human tolerance<br />

4. Habitat fragmentation (result<strong>in</strong>g <strong>in</strong> non-viable pops)<br />

Europe<br />

[15]<br />

Least Concern Increas<strong>in</strong>g Large size<br />

Increas<strong>in</strong>g<br />

As above<br />

Europe –<br />

regional<br />

populations<br />

[15]<br />

Critically<br />

Endangered:<br />

Central Europe,<br />

Sierra Morena<br />

Endangered:<br />

Westerncentral<br />

Alps,<br />

Scand<strong>in</strong>avia<br />

Increas<strong>in</strong>g:<br />

Iberia, Westerncentral<br />

Alps, Italian<br />

pen<strong>in</strong>sula<br />

Decreas<strong>in</strong>g: Karelia<br />

Small size<br />

1. Human persecution<br />

2. Lack/fragmentation of management regimes<br />

3. Small population size<br />

4. Habitat fragmentation<br />

5. Low genetic variability<br />

Vulnerable:<br />

Italian<br />

pen<strong>in</strong>sula<br />

Near<br />

Threatened:<br />

Iberia, Karelia<br />

Least Concern:<br />

D<strong>in</strong>aric-Balkan,<br />

Carpathian,<br />

Baltic<br />

97

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