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130925-studie-wildlife-comeback-in-europe

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% change<br />

20,000<br />

16,000<br />

12,000<br />

200<br />

100<br />

0<br />

Figure 2.<br />

Change <strong>in</strong> Eurasian<br />

beaver population<br />

abundance by<br />

decade and overall<br />

change between<br />

1960 and 2005. Please<br />

note that due to the<br />

way change was<br />

calculated, decadal<br />

change does not sum<br />

to overall change.<br />

1960s 1970s 1980s 1990s 2000–05 1960–2005<br />

Drivers of recovery<br />

Although none of the factors tested expla<strong>in</strong>ed the<br />

resurgence of Eurasian beaver observed <strong>in</strong> our<br />

data set, a number of possible reasons for these<br />

trends have been discussed <strong>in</strong> the literature. The<br />

most important of these are undoubtedly legal<br />

protection and the restriction of hunt<strong>in</strong>g, re<strong>in</strong>troductions<br />

and translocations, and natural recolonisation<br />

follow<strong>in</strong>g <strong>in</strong>itial recovery (Table 3).<br />

Legal protection of the five populations<br />

rema<strong>in</strong><strong>in</strong>g at the beg<strong>in</strong>n<strong>in</strong>g of the 20 th century [5]<br />

was key <strong>in</strong> enabl<strong>in</strong>g the species to persist <strong>in</strong> Europe.<br />

In addition to merely preserv<strong>in</strong>g genetic material,<br />

some of these f<strong>in</strong>al five also acted as source populations<br />

for many of the extensive re<strong>in</strong>troduction<br />

and translocation programmes that have taken<br />

place <strong>in</strong> at least 25 European countries [5] . Initially,<br />

these efforts were motivated by the fur trade,<br />

comprised hard releases and lacked habitat suitability<br />

assessments, but latterly they became more<br />

conservation-focused and better researched [5] .<br />

Viable populations have been established <strong>in</strong> all<br />

countries where re<strong>in</strong>troductions have occurred [5]<br />

because the species is robust and will succeed <strong>in</strong><br />

most river systems if left alone [2] . As part of legal<br />

protection, the cessation of hunt<strong>in</strong>g, which had<br />

driven the species to near ext<strong>in</strong>ction, has resulted<br />

<strong>in</strong> population growth, e.g. <strong>in</strong> Belarus [5] .<br />

A robust species, the beaver expands rapidly<br />

with<strong>in</strong> a watershed of occurrence to occupy<br />

available optimum habitat [20] . However, <strong>in</strong> medium<br />

to large-sized watersheds [2] this reduces beaver<br />

density, and the critical threshold for encounter<strong>in</strong>g<br />

a mate is usually only reached after 10–25<br />

years, lead<strong>in</strong>g to an explosion <strong>in</strong> numbers [20] . In<br />

smaller rivers, for example the River Tay <strong>in</strong> the<br />

United K<strong>in</strong>gdom, this phase of rapid <strong>in</strong>crease<br />

occurs almost immediately [2] . The follow<strong>in</strong>g 10<br />

or so years are marked by population decl<strong>in</strong>e as<br />

marg<strong>in</strong>al habitats become exhausted, and this is<br />

followed by rough stability [20] . While the species<br />

can easily spread between watersheds – as has<br />

been the case <strong>in</strong> eastern Norway, which was repopulated<br />

from Sweden, the islands of Saaremaa and<br />

Hiiumaa from ma<strong>in</strong>land Estonia [5] , and Slovenia<br />

from Croatian watersheds [2] – natural and artificial<br />

barriers can strongly h<strong>in</strong>der expansion [5] .<br />

While legal protection, re<strong>in</strong>troductions and<br />

recolonisation are the key drivers of beaver<br />

recovery <strong>in</strong> Europe, habitat protection and restoration<br />

have also played a role <strong>in</strong> its resurgence.<br />

Unsuitable habitat is believed to be the reason for<br />

re<strong>in</strong>troduction failures <strong>in</strong> Switzerland, and poor<br />

habitat quality is limit<strong>in</strong>g reproductive output <strong>in</strong><br />

the Biesbosch National Park <strong>in</strong> the Netherlands [5] .<br />

On the other hand, the conservation and regeneration<br />

of riparian zones around rivers for flood<br />

control has created suitable beaver habitat around<br />

the cont<strong>in</strong>ent [5] , and has certa<strong>in</strong>ly contributed to<br />

the observed recoveries. In addition to changes<br />

<strong>in</strong> the way that man is manag<strong>in</strong>g the Eurasian<br />

beaver, the species’ resilience and ability to spread<br />

swiftly with<strong>in</strong> a watershed follow<strong>in</strong>g <strong>in</strong>troduction<br />

or colonisation, will also have been beneficial <strong>in</strong><br />

the <strong>in</strong>crease and expansion observed throughout<br />

most of its historic range. Populations have now<br />

been established <strong>in</strong> all former range countries<br />

except Portugal, Italy and the southern Balkans [11] .<br />

Recent developments<br />

Despite the impressive recovery <strong>in</strong> abundance<br />

and range, the Eurasian beaver still occurs at<br />

low densities <strong>in</strong> some locations, and large areas<br />

of suitable habitat rema<strong>in</strong> unused [11] , suggest<strong>in</strong>g<br />

plenty of opportunity for further spread. Considerable<br />

growth <strong>in</strong> range and numbers is expected,<br />

particularly <strong>in</strong> western Europe and the Danube<br />

watershed [11] . This would make the species<br />

common <strong>in</strong> much of Europe with<strong>in</strong> the next few<br />

decades [10] . In addition, re<strong>in</strong>troduction efforts<br />

are cont<strong>in</strong>u<strong>in</strong>g with an ambitious programme to<br />

recolonise the entire Danube bas<strong>in</strong>, with other<br />

successful recent re<strong>in</strong>troductions (or spread) <strong>in</strong><br />

Denmark, Belgium and Scotland [21] . As a result,<br />

many populations are now considered to be at<br />

a mature stage of development [5] . There are also<br />

proposals to re-establish the beaver <strong>in</strong> Wales [7]<br />

and England [22] . The species is expected to spread<br />

from the Tay watershed <strong>in</strong> Scotland, where the<br />

population was discovered <strong>in</strong> 2010 and <strong>in</strong>creased<br />

by around 20% s<strong>in</strong>ce 2012, to other watersheds<br />

such as the Dee, Spey and Forth <strong>in</strong> the United<br />

K<strong>in</strong>gdom [2] . Although the population was established<br />

“illegally”, it is unlikely that there will be<br />

the resources and public support to eradicate<br />

it [2] . In Luxembourg, newly established beavers<br />

were found to be North American escapees from<br />

154

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