130925-studie-wildlife-comeback-in-europe
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% change<br />
20,000<br />
16,000<br />
12,000<br />
200<br />
100<br />
0<br />
Figure 2.<br />
Change <strong>in</strong> Eurasian<br />
beaver population<br />
abundance by<br />
decade and overall<br />
change between<br />
1960 and 2005. Please<br />
note that due to the<br />
way change was<br />
calculated, decadal<br />
change does not sum<br />
to overall change.<br />
1960s 1970s 1980s 1990s 2000–05 1960–2005<br />
Drivers of recovery<br />
Although none of the factors tested expla<strong>in</strong>ed the<br />
resurgence of Eurasian beaver observed <strong>in</strong> our<br />
data set, a number of possible reasons for these<br />
trends have been discussed <strong>in</strong> the literature. The<br />
most important of these are undoubtedly legal<br />
protection and the restriction of hunt<strong>in</strong>g, re<strong>in</strong>troductions<br />
and translocations, and natural recolonisation<br />
follow<strong>in</strong>g <strong>in</strong>itial recovery (Table 3).<br />
Legal protection of the five populations<br />
rema<strong>in</strong><strong>in</strong>g at the beg<strong>in</strong>n<strong>in</strong>g of the 20 th century [5]<br />
was key <strong>in</strong> enabl<strong>in</strong>g the species to persist <strong>in</strong> Europe.<br />
In addition to merely preserv<strong>in</strong>g genetic material,<br />
some of these f<strong>in</strong>al five also acted as source populations<br />
for many of the extensive re<strong>in</strong>troduction<br />
and translocation programmes that have taken<br />
place <strong>in</strong> at least 25 European countries [5] . Initially,<br />
these efforts were motivated by the fur trade,<br />
comprised hard releases and lacked habitat suitability<br />
assessments, but latterly they became more<br />
conservation-focused and better researched [5] .<br />
Viable populations have been established <strong>in</strong> all<br />
countries where re<strong>in</strong>troductions have occurred [5]<br />
because the species is robust and will succeed <strong>in</strong><br />
most river systems if left alone [2] . As part of legal<br />
protection, the cessation of hunt<strong>in</strong>g, which had<br />
driven the species to near ext<strong>in</strong>ction, has resulted<br />
<strong>in</strong> population growth, e.g. <strong>in</strong> Belarus [5] .<br />
A robust species, the beaver expands rapidly<br />
with<strong>in</strong> a watershed of occurrence to occupy<br />
available optimum habitat [20] . However, <strong>in</strong> medium<br />
to large-sized watersheds [2] this reduces beaver<br />
density, and the critical threshold for encounter<strong>in</strong>g<br />
a mate is usually only reached after 10–25<br />
years, lead<strong>in</strong>g to an explosion <strong>in</strong> numbers [20] . In<br />
smaller rivers, for example the River Tay <strong>in</strong> the<br />
United K<strong>in</strong>gdom, this phase of rapid <strong>in</strong>crease<br />
occurs almost immediately [2] . The follow<strong>in</strong>g 10<br />
or so years are marked by population decl<strong>in</strong>e as<br />
marg<strong>in</strong>al habitats become exhausted, and this is<br />
followed by rough stability [20] . While the species<br />
can easily spread between watersheds – as has<br />
been the case <strong>in</strong> eastern Norway, which was repopulated<br />
from Sweden, the islands of Saaremaa and<br />
Hiiumaa from ma<strong>in</strong>land Estonia [5] , and Slovenia<br />
from Croatian watersheds [2] – natural and artificial<br />
barriers can strongly h<strong>in</strong>der expansion [5] .<br />
While legal protection, re<strong>in</strong>troductions and<br />
recolonisation are the key drivers of beaver<br />
recovery <strong>in</strong> Europe, habitat protection and restoration<br />
have also played a role <strong>in</strong> its resurgence.<br />
Unsuitable habitat is believed to be the reason for<br />
re<strong>in</strong>troduction failures <strong>in</strong> Switzerland, and poor<br />
habitat quality is limit<strong>in</strong>g reproductive output <strong>in</strong><br />
the Biesbosch National Park <strong>in</strong> the Netherlands [5] .<br />
On the other hand, the conservation and regeneration<br />
of riparian zones around rivers for flood<br />
control has created suitable beaver habitat around<br />
the cont<strong>in</strong>ent [5] , and has certa<strong>in</strong>ly contributed to<br />
the observed recoveries. In addition to changes<br />
<strong>in</strong> the way that man is manag<strong>in</strong>g the Eurasian<br />
beaver, the species’ resilience and ability to spread<br />
swiftly with<strong>in</strong> a watershed follow<strong>in</strong>g <strong>in</strong>troduction<br />
or colonisation, will also have been beneficial <strong>in</strong><br />
the <strong>in</strong>crease and expansion observed throughout<br />
most of its historic range. Populations have now<br />
been established <strong>in</strong> all former range countries<br />
except Portugal, Italy and the southern Balkans [11] .<br />
Recent developments<br />
Despite the impressive recovery <strong>in</strong> abundance<br />
and range, the Eurasian beaver still occurs at<br />
low densities <strong>in</strong> some locations, and large areas<br />
of suitable habitat rema<strong>in</strong> unused [11] , suggest<strong>in</strong>g<br />
plenty of opportunity for further spread. Considerable<br />
growth <strong>in</strong> range and numbers is expected,<br />
particularly <strong>in</strong> western Europe and the Danube<br />
watershed [11] . This would make the species<br />
common <strong>in</strong> much of Europe with<strong>in</strong> the next few<br />
decades [10] . In addition, re<strong>in</strong>troduction efforts<br />
are cont<strong>in</strong>u<strong>in</strong>g with an ambitious programme to<br />
recolonise the entire Danube bas<strong>in</strong>, with other<br />
successful recent re<strong>in</strong>troductions (or spread) <strong>in</strong><br />
Denmark, Belgium and Scotland [21] . As a result,<br />
many populations are now considered to be at<br />
a mature stage of development [5] . There are also<br />
proposals to re-establish the beaver <strong>in</strong> Wales [7]<br />
and England [22] . The species is expected to spread<br />
from the Tay watershed <strong>in</strong> Scotland, where the<br />
population was discovered <strong>in</strong> 2010 and <strong>in</strong>creased<br />
by around 20% s<strong>in</strong>ce 2012, to other watersheds<br />
such as the Dee, Spey and Forth <strong>in</strong> the United<br />
K<strong>in</strong>gdom [2] . Although the population was established<br />
“illegally”, it is unlikely that there will be<br />
the resources and public support to eradicate<br />
it [2] . In Luxembourg, newly established beavers<br />
were found to be North American escapees from<br />
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