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from tourism [15] and it is reasonable to <strong>in</strong>fer that<br />

the draw of <strong>wildlife</strong> has been beh<strong>in</strong>d much of the<br />

<strong>in</strong>crease <strong>in</strong> tourism <strong>in</strong> similar areas of high biodiversity.<br />

For example <strong>in</strong> South Africa where safari<br />

opportunities are a major attraction, tourism<br />

accounts for 7.7% of GDP [18] and <strong>in</strong> Namibia, the<br />

communal conservancy scheme implemented<br />

by the government has generated an <strong>in</strong>creas<strong>in</strong>g<br />

contribution to the national economy between<br />

1990 and 2011 [19] .<br />

Other than direct payments <strong>in</strong> the form of<br />

entrance fees to national parks, costs for a tour<br />

guide and accommodation, <strong>in</strong>direct payments<br />

can also be important <strong>in</strong> support<strong>in</strong>g the local<br />

and national economy through the purchase of<br />

products and services from other sectors both<br />

by tourists and by residents whom the tourist<br />

<strong>in</strong>come has benefitted [15] . Provided that certa<strong>in</strong><br />

measures are <strong>in</strong> place, the <strong>in</strong>flux of visitors to view<br />

<strong>wildlife</strong> can provide economic relief to the often<br />

poor rural communities [15] . The TAMAR project <strong>in</strong><br />

Brazil is one such enterprise which has achieved<br />

success <strong>in</strong> attract<strong>in</strong>g tourists to visit mar<strong>in</strong>e turtle<br />

nest<strong>in</strong>g sites, result<strong>in</strong>g <strong>in</strong> both socio-economic<br />

and conservation benefits [20] . The project <strong>in</strong>corporates<br />

the tra<strong>in</strong><strong>in</strong>g and employment of local people,<br />

extend<strong>in</strong>g the benefits from visitors to the wider<br />

community, and implement<strong>in</strong>g mar<strong>in</strong>e turtle<br />

research for conservation [15] . However meet<strong>in</strong>g the<br />

demand of the tourist <strong>in</strong>dustry can often mean<br />

rely<strong>in</strong>g on foreign <strong>in</strong>vestment and <strong>in</strong> some cases,<br />

part of the economic benefit is appropriated to<br />

other countries [15] .<br />

In more developed regions, there has long<br />

been an <strong>in</strong>terest <strong>in</strong> <strong>wildlife</strong> related activities and<br />

this sector is fairly well established. Prelim<strong>in</strong>ary<br />

f<strong>in</strong>d<strong>in</strong>gs <strong>in</strong> the USA suggest that hunt<strong>in</strong>g, fish<strong>in</strong>g<br />

and <strong>wildlife</strong> watch<strong>in</strong>g for recreation accounted for<br />

1% of national GDP <strong>in</strong> 2011 and such activities are<br />

considered pivotal for local and national economic<br />

growth [21] . Wildlife watch<strong>in</strong>g was the most popular<br />

activity, more popular than sport fish<strong>in</strong>g and<br />

hunt<strong>in</strong>g comb<strong>in</strong>ed. At the EU level, a 2011 study for<br />

the Commission estimated the economic value of<br />

the benefits provided by tourism, recreation and<br />

employment supported by Natura 2000, a network<br />

of nearly 26,000 protected sites <strong>in</strong> the EU. The study<br />

suggested that expenditure related to tourism and<br />

recreation supported by Natura 2000 sites was<br />

between €50 and €85 billion <strong>in</strong> 2006, support<strong>in</strong>g<br />

between 4.5 and 8 million full-time-equivalent<br />

jobs [22] .<br />

At the national level, there is a considerable<br />

economic opportunity for <strong>wildlife</strong> related activities<br />

with an exist<strong>in</strong>g and <strong>in</strong>terested market. In the<br />

UK for example, over half of the adult population<br />

of England visit the natural environment at least<br />

once a week [23] . In Scotland, no less than 56% of<br />

all travel is nature oriented [24] and it is hav<strong>in</strong>g<br />

clear economic benefits. Work commissioned by<br />

Scottish Natural Heritage estimated that total<br />

annual visitor spend<strong>in</strong>g attributable to naturebased<br />

tourism (<strong>in</strong>clud<strong>in</strong>g outdoor activities other<br />

than <strong>wildlife</strong> watch<strong>in</strong>g) is £1.4 billion with 39,000<br />

associated full-time-equivalent jobs [25] . The opportunity<br />

for real economic benefits from <strong>wildlife</strong><br />

resurgence, and the nature tourism that can<br />

be generated from that, seems to be realistic <strong>in</strong><br />

several areas of Europe.<br />

The presence of a key charismatic species can<br />

act as a great <strong>in</strong>centive for people to take part <strong>in</strong><br />

<strong>wildlife</strong> watch<strong>in</strong>g and sometimes to pay a lot of<br />

money for the privilege. For <strong>in</strong>stance, a permit to<br />

see the Mounta<strong>in</strong> gorillas (Gorilla ber<strong>in</strong>gei) of the<br />

Virunga mounta<strong>in</strong> range of Rwanda, Democratic<br />

Republic of Congo and Uganda costs each <strong>in</strong>ternational<br />

tourist between $400 and $750 [26] , which<br />

has the benefit of generat<strong>in</strong>g local and national<br />

revenue. In addition, the presence of regular<br />

visitors and monitor<strong>in</strong>g is one of the reasons<br />

contribut<strong>in</strong>g towards the survival and potential<br />

recovery of this threatened population [27] . There<br />

are issues surround<strong>in</strong>g revenue-shar<strong>in</strong>g amongst<br />

all stakeholders but whilst there are still challenges<br />

to be overcome [28] , this example shows that there<br />

are <strong>in</strong>dividual species that provide enough of a<br />

draw to attract high numbers of visitors.<br />

Similarly, the recovery of charismatic species<br />

<strong>in</strong> Europe could prove to be an economic opportunity<br />

as well as an ecological one. As the Abruzzo<br />

region of Italy started be<strong>in</strong>g marketed as bear and<br />

wolf country, due to recent resurgence <strong>in</strong> populations,<br />

the number of tourists to the area <strong>in</strong>creased<br />

even though the likelihood of actually see<strong>in</strong>g<br />

any <strong>in</strong>dividuals of those species is still low [29] . The<br />

Brown bear (Ursus arctos) is also the flagship species<br />

for the Somiedo Natural Park <strong>in</strong> northern Spa<strong>in</strong><br />

and has attracted <strong>in</strong>creas<strong>in</strong>g numbers of tourists<br />

to the local area almost exclusively due to the use<br />

of the bear as a means for promot<strong>in</strong>g tourism [30] .<br />

An assessment of <strong>wildlife</strong> watch<strong>in</strong>g <strong>in</strong> F<strong>in</strong>land<br />

suggests that the number of visitors <strong>in</strong>creased by<br />

90% between 2005 and 2008 with the majority of<br />

tourists drawn by the presence of predators such as<br />

Brown bear and Wolver<strong>in</strong>e [31] . In economic terms,<br />

prelim<strong>in</strong>ary results estimated a turnover from<br />

this sector of €4–5 million <strong>in</strong> 2012 with <strong>wildlife</strong><br />

watch<strong>in</strong>g and photography as the ma<strong>in</strong> reason for<br />

an estimated 73% of tourists who visited F<strong>in</strong>land [32] .<br />

In Cevennes National Park <strong>in</strong> France, the re<strong>in</strong>troduced<br />

Griffon (Gyps fulvus) and C<strong>in</strong>ereous vultures<br />

(Aegypius monachus) have provided new bird<br />

watch<strong>in</strong>g opportunities for tourists, attract<strong>in</strong>g<br />

80,000 visitors per year. It was also estimated that<br />

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