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The genus Cinnamomum

The genus Cinnamomum

The genus Cinnamomum

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Botany and Crop Improvement of Cinnamon and Cassia 51<br />

divisions producing two superposed tiers of cells (pc and pc). <strong>The</strong>y undergo further<br />

divisions and finally lead to the formation of the central cylinder of the stem and cotyledons<br />

and the hypocotyledonary region, respectively. <strong>The</strong> two daughter cells of the basal<br />

cell (bc) divide transversely giving rise to two superposed tiers of cells. <strong>The</strong> upper tier<br />

divides further in all planes and forms the hypophyseal region. <strong>The</strong> lower tier gives rise<br />

to the suspensor. <strong>The</strong> epiphyseal cell undergoes further divisions, ultimately forming<br />

the stem apex.<br />

Since the derivatives of the apical cell of the two-celled proembryo alone contribute<br />

to the formation of the embryo proper, while the suspensor and hypophyseal regions are<br />

derived from the basal cell, the embryo development is of Onagraceae type.<br />

Endosperm<br />

ca<br />

cb<br />

1 2 3<br />

e<br />

7<br />

10<br />

e<br />

e<br />

e<br />

pc<br />

pc′<br />

e<br />

4<br />

8 9<br />

11<br />

m<br />

ei<br />

e e<br />

12<br />

5 6<br />

Figure 2.16 Stages in the development of embryo (for explanations see text). ca – apical cell; cb –<br />

basal cell; e – epiphyseal initial; pc, pc – tiers of cells derived from subepiphyseal cells;<br />

m – hypophyseal initial, that divides and forms the hypophyseal region; ci – suspensor<br />

initial that eventually gives rise to the suspensor. (Source: Sastri, 1958.)<br />

<strong>The</strong> primary endosperm nucleus divides earlier than the fertilised egg. Endosperm<br />

development is of the nuclear type (Sastri, 1958). By the time the zygote undergoes the<br />

first division, eight to ten free nuclei derived from the division of the endosperm nucleus<br />

are seen distributed along the periphery of the embryosac. <strong>The</strong>se nuclei are large with<br />

one to three nucleoli. <strong>The</strong> endosperm nuclei have glistening crystalline bodies at the<br />

13

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