ENVIRONMENTAL CONSEQUENCES in rocky mountain coniferous ...

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Recently Harvey and others (in process), through 1 4 analysis, ~ have shown that Douglas-fir (~seudotsup menziesii (Mirb. 1 Franco) residues persist for 500 years or longer, depending on the site and depth in soil. Our observations, though incomplete, indicate that brown-rotted Douglas-fir residue is the most persistent, We tentatively suggest that resistance to further decomposition is a function of extractives or extractive-1 i ke substances chemically bound to the I ignin polymer in Douglas-fir--to a greater extent than other conifer species. Appearance and characteristics of brown-cubical decayed wood The substrate is readily characterized in the field by its dark brown to reddish-brown color and its texture is three-dimensional 1 v checked, forming sixsided configurations (often referred to as cubical ) (fig .- 9) . Figure 9. -- Brown-rotted L@ugZas-ffr shoving fissures and in the subs*rate. checks
Recognizable brown-cubical decayed wood in the field is usually in the form of dead fallen trees or logging residues greater than 0.5 m in diameter or large enough to maintain a reasonable degree of recognizability through the processes of decay, weathering, and physical dispersion. Residues of lesser dimension are subject to spreading, dispersion, and downhill movement in steep terrain. Thus, the larger residues tend to collapse and spread during the deterioration process, appearing somewhat ellipsoidal in profile at the soil surface. Tabl es 3 and 4 summarize the more important properties and characteristics of soil wood. Ryp'aFek and Ryp'ac'kova (1975) note that humic substances isolated from wood decomposed by brown rot fungi exhibit properities sSmi1ar to those of soil humus, and that dif- ferences are dependent on the degree of wood decomposition. Tabl e 3 .--Summary of brown-cubical decayed soil wood properties and characteristics. Volume 5-25 pct 0.5-0.8 KgN/ha/YR - N2 Fixation Persistence 500 + 12-fold increase in N compared to un- decayed wood Moisture 300 pct + 5-fold increase in P CEC 45-90-MEQ Appreciabl e niacroel ement content Per 100 gm Low pH 4-5.5 Lignin content 68 pct Tabl e 4.-- Mean percent 1 ignin, ash and carbohydrate contents, based on odw of brown cubical decayed wood from Coram Experimental Forest, Montana, 1975. --- Number Ash Ligni n Carbohydrate sampl es content content content ........................................................................... " NONDECAY" ACTIVITIES In addition to the two major kinds of decay associated with biodegradation of woody residues, other activities of fungi may be considered nondegradative in the sense that they are not responsible for major volume or weight reductions in residues. Ecologically, these fungi appear early in the biodegradative sequence of plant materials, They are the primary colonizers with the ability to metabolize the readily available low-molecular-weight carbon and nitrogen sources that are ' available in fresh residues. Fungal numbers gradually decrease with time, an increase in extractive content and, perhaps, increasing acidity of substrates. Their abil i ty to rapidly form large amounts of biomass within short periods of time is noticeable in figures 10 and 11.
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ENVIRONMENTAL CONSEQUENCES OF TIMBE
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USDA Forest Service General Technic
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REW 'ORD One of the pressing proble
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BIOLOGICAL IMPLICATIONS Biological
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By the mid-1 960's the apparently u
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Scientist, audience: "The Three-Mil
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I believe that the future of forest
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enewable resource that can be proce
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productive capacity. Recent legisla
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Much of the research investigating
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Six sale area blocks were logged (f
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Understory Removal Figure 4.--Eccpe
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Gentle slopes and easy access to cu
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Each of the four cutting units was
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A concerted effort was made to coor
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oriented toward the basic response
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WOODY MATERIAL IN NORTHERN ROCKY MO
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STANDING, 3 " 1 DIA. ? (GREEN & DEA
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I 11 Table 1,--Volume of wood by co
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Small material under 3 inch (7.6 cm
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Depending on the cutting and treatm
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MICROENVIRONMENTAL RESPONSE TO HARV
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I Where: Cs is the concentration at
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Figure I.-- Residue treatments used
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HARVESTING AND RESIDUE INFLUENCES R
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I 1 1 1 1 1 1 1 1 1 1 1 JAN. FEB. M
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Sensible heat flux, evaporative flu
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SOLO, 1977- 78 UNCUT CUT Mean max.
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It is logical that if surface tempe
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0 - 5 - 10 - 15 20 - Chips - Clrare
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Brown, crumbly, decayed material (B
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At Coram, steep harvested slopes ha
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surface of unburned and hard-burned
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canopy. Light then indirectly cause
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Moisture Moisture is frequently ide
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Net radiation--the measure of the a
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Table 5.--Possible solutions for po
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Cochran, P. H. 1969. Thermal proper
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Lommen, P. W., Cr, R. Schwintzer, C
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Stark, Nel 1 ie. 1980. The impacts
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and water use during the growing se
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The main study site (fig. 1) occupi
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The four residues util i zation tre
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RAIN The major precipitation instru
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Figure 5.--SG,L. wuwr zc)as measure
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S treamf 1 ow A &foot H-flume was i
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Figure 7.--Snow water equivalent on
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Soi 1 Water Water present in the so
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SILVICULTURE TREATMENTS Si 1 vicul
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As indicated in figure 11 and appen
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The most consistent difference in w
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MEAN CONSUMPTIVE USE -ALL YEARS (19
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Lowest daily water use occurred at
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6. Water use began in Apri 1 , acce
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Klages, M. G., R. C. McConnell , an
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APPENDICES Appendix A.--Hydrograph
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Appendix C.--Rainfall reaching the
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Appendix E.--Water use during the g
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One of the most efficient methods o
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Control Chipped Picked up Broadcast
Page 121 and 122: Control Chapped Picked up Broadcast
Page 123 and 124: Control Chipped Picked up Broadcast
Page 125 and 126: ' Control Chipped and Picked up and
Page 127 and 128: One of the least detrimental residu
Page 129 and 130: INTRODUCTION In the early days, the
Page 131 and 132: %pire 1.--Measured concentrations (
Page 133 and 134: The silvicul tural treatments appli
Page 135 and 136: I 2 Table 2. --Percent of total qua
Page 137 and 138: Unmerchantable TOP Shrubs Lower Cro
Page 139 and 140: create a nitrogen deficiency during
Page 141 and 142: Steele, R. W. 1975. Understory burn
Page 143 and 144: National Forest. This paper summari
Page 145 and 146: UNIT 2 CONVENT IONALLY Du Bois, Wyo
Page 147 and 148: Trees Figures 3 and 4 show the oven
Page 149 and 150: Weights of typical trees on these t
Page 151 and 152: pi1 ed-burned treatments were lowes
Page 153 and 154: MINERAL SOIL Total nutrient content
Page 155 and 156: Tab1 e 5. --Concentrations of avai
Page 157 and 158: Table 6. --Average concentrations o
Page 159 and 160: sites. Burning then transformed the
Page 161 and 162: RESIDUE DECAY PROCESSES AND ASSOCIA
Page 163 and 164: The hardwood or angiospermous timbe
Page 165 and 166: Woody Substrates Gyrnnospermsr (Int
Page 167 and 168: Natural inoculation of residues may
Page 169 and 170: On the Coram site, brown-rot fungi
Page 171: Table 2.--Probability (P) of encoun
Page 175 and 176: I Newly formed, small-dimension res
Page 177 and 178: Highley, T. L. 1976. Hemicellulases
Page 179 and 180: MICROBIAL PROCESSES ASSOCIATED WITH
Page 181 and 182: Nonsymbiotic N Fixation Free-1 ivin
Page 183 and 184: In a previous paper (Jurgensen and
Page 185 and 186: Symbiotic N Fixation The establishm
Page 187 and 188: carbon and NHq (Ahlgren, 1974). Mic
Page 189 and 190: Prescribed f~re CLEARCUT - BURN CLE
Page 191 and 192: Hungerford, R.9. 1980. Microenvi ro
Page 193 and 194: ECOLOGY OF ECTOMYCORRHIZAE IN NORTH
Page 195 and 196: Coram - Subalpine fir Site (ABLAICL
Page 197 and 198: Figure 2.-- ReZative yield capabiZi
Page 199 and 200: Figure 5. -- Percentage of soCZ-woo
Page 201 and 202: Figure 9,-- Percentage of toid ectm
Page 203 and 204: Effect of Soil Components on Ectomy
Page 205 and 206: Effect of Organic Matter Quantity o
Page 207 and 208: PARTIAL CUT Effect of Harvesting on
Page 209 and 210: Figure 20. -- Average nwnbers of ac
Page 211 and 212: Gijbl , F. 1967. Mykorrhizauntersuc
Page 213 and 214: BIOLOGICAL IMPLICATIONS Initial cha
Page 215 and 216: FOREST SOIL BIOLOGY The act of incr
Page 217 and 218: the soil will retain its ability to
Page 219 and 220: Since the best quantity of organic
Page 221 and 222: We can also visualize situations on
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Harvey, A. E., M. F. Jurgensen, and
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INTRODUCTION Vegetation integrates
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Because of cool, wet weather in 197
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We determined volume and cover usin
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The stems were divided at 4 mm (0.1
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RESIDUES TREATMENT EFFECTS As descr
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Overall shrub recovery rates do not
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Rose behaved similar to ninebark (f
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Meanwhile, small shrub cover change
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Biomass Regression analyses relatin
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SILVICULTURE TREATMENT EFFECTS Harv
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RESIDUES TREATMENT EFFECTS Major sh
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Lesser vegetation--herbs and small
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Leaf and stem components of the shr
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APPENDIX I Trees and shrubs found o
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INTRODUCTION Many western larch (La
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Figure 2. --Lower cutting blocks on
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each permanent point) of the clearc
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Figure 3.--Marking gemination and c
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were not significant. These treatme
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Tab1 e 2. --Fi 11 ed conifer seed (
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Dispersal of sound seed on the uppe
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Table $.--Number of 3- and 5-year-o
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Table 6. --Germination of conifers
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Survival, growth and form will be m
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Es tab1 ishment and Initial Develop
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to the amount of residue originally
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As the following tabulation shows,
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Soot-Seeded Broadcast burning, and
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Natural Regeneration Natural regene
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.4s shown in fig. 5, vegetation was
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Interestingly, the high phenol leve
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DeByle, Norbert V. 1980. Harvesting
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EFFECT OF SILVICULTURAL PRACTICES,
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mountain pine beetle. These two for
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GROUP SELECTION 22 CONTROL SHELTERW
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FIELD AND LABORATORY PROCEDURES To
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I Many of the groups were trapped v
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shelterwood with residues, were sig
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HARVESTING AND RESIDUE MANAGEMENT T
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indicate a treatment effect of burn
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Undisturbed Forest .". . . . Clearc
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1 equally abundant in all treatment
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I not seem to be directly caused by
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the undisturbed forests. Some resea
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Huhta, V., E. Karppinen, M. Nurmine
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POPULATIONS OF SOME FOREST LITTER,
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STUDY DESIGN The study area is loca
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GROUP SELECTION 22 CONTROL '... . .
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Sampling in 1977 was restricted to
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Table 1 .--Mean total mesofauna pop
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0 - (JUNE) from S helterwood (AUGUS
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SHELTERWOOD--RESIDUE BURNED The eff
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In the first season or two followin
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LITERATURE CITED Ahlgren, I. F. 197
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A REVIEW OF SOME INTERACTIONS BETWE
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and Keen 1960; Felix and others 197
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Pissodes strobi Peck, In a thinned
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foresters feel as though "It's 1 i
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The large aerial spray programs aga
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Weakened living trees, or trees kil
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Most western pine beetle attacks in
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The galleries, or mines, of wood bo
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Other wood borers. --Several specie
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Deterioration of spruce (Picea a. )
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The eruption of Mount Saint Helens
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Bark and engraver beetles infesting
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In the northern Rockies, one conife
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management, and not the beetle." If
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investigation regardin controlled b
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WOOD BORERS Mitchell and Martin (In
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Still, by 1938, there was a feeling
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I Different i nvesti gators have us
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if the forest floor is completely c
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Carabids as biological control agen
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een placed for protection from rode
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with every tree in the stand contin
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Spiders as predators.--Studies have
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At least two studies have been made
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In the long-leaf pine forests in th
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RESIDUES, FIRES, INSECTS, AND FORES
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In the spruce-fir stands infested w
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Recently, Senator John Me1 cher (Mo
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plans. Since fire, too, is a decomp
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CHANGING FOREST INSECT PROBLEMS Ins
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Basham, J. T. 1957. The deteriorati
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Cole, D. M. 1978. Feasibility of si
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Fellin, D. G. and P. C. Johnson. 19
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Graham, S. A. 1922. Some entomoloqi
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Huhta, V., M. Nurminen and A. Valpa
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Lawrence, W. H. and 3. H. Rediske.
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Miller, J. M. and J. E. Patterson.
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Rice, L * A. 1932. The effect of fi
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Stoddard, H. L. Sr. 1963. Bird habi
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Weaver, Harol d. 1951. Fire as an e
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RESOURCE MANAGEMENT IMPLICATIONS Th
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INFLUENCE OF HARVESTING AND RESIDUE
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F
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NEAR COMPLETE Figure 2.--Byrmnts f
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Figure 3.-- Bymrmrs fireZine intens
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F5qure 4. --Byramrs fireZ-Cne inten
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1. Nomographs of Rate of Spread, Fi
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3. Consider other fire-related fact
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Puckett, John V., Cameron M. Johnst
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Important and rapid progress has be
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tlarvested Areas What happened when
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UNCUT SHELTERWOOD Protect understor
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Initially the areas are rated low.
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The primary reason for this discrep
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f n contrast to displaced bears, so
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Quantitative data on grizzly behavi
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Asherin, Duane A. 1976. Changes in
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Leege, Thomas A. 1969. Burning sera
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Reynolds, Hudson G. 1969. Aspen gro
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INFLUENCES OF HARVESTING AND RESIDU
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CLEARCUT LOGGING & COMPLETE BURN DE
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C LEARCUT LOGGING & BROADCAST BURN
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EFFECTS OF SMALL MAMMALS ON FOREST
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I I Habitat Manipulation Small mamm
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Fala, Robert A. 1975. Effects of pr
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Tevis, Lloyd Jr. l956b. Pocket goph
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forests are cavity nesters. They ar
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GROUP SELECTION: A1 1 merchantab le
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TABLE 3. Pre- and post-logging volu
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Censuses Censuses for all species (
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Fbgure 6.-- Lightn
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Pileated Woodpeckers on the CEF nes
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TABLE 7. Percent of sampling time t
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I Observed woodpecker feeding Down,
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Table 9 shows feeding time in the l
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L c - 10- ul 2 0 8- 6 - 4 2 - - 0 .
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Figure 15. -- A Hairy Woodpecker ne
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~
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oth offered advice on study site se
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Hairy Woodpecker Downy Woodpecker B
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THE SITUATION Forest land managers
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In the descriptive approach, howeve
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ORGANIZING INFORMATION One differen
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As has already been stated, the for
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Figure 7.--Matrix of interactions b
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understory biomass density, X2 repr
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or, by rearrangement, AX. I afi Ax,
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DISCUSSION It is important to note
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LITERATURE CITED Billings, W. D. 19
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Richard T. Wick Burlington Northern
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themselves, the traditional industr
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highways. Of course, all roads are
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Darrel L. Kenops District Ranger US
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In the future we will see our const
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Lyon, L. Jack, Project Leader, Ecol
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The Intermountain Station, headquar
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