ENVIRONMENTAL CONSEQUENCES in rocky mountain coniferous ...

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infra-red radiation for a distance of several miles through paired sense organs on the mesothorax (Evans 1964; Boag and Evans 1967); the organs are so sensitive that extraneous radiation from sources outside of the temperature range of wi 1 dfi res is effectively filtered (Evans 1966). This feature allows Melanophila to be among the first insects to reach fire-killed trees--usually before competitors--and has resulted in the Holarctic distribution of at least one species, M. acuminata (Evans 1971). Buprestids of the genus Melanophila are known in some areas of North Carol ina as "fire bugs," and have been observed landing on stumps that were still glowing (Linsley 1943). Another flatheaded borer, Asemum atrum Esch., infested the sapwood of the majority of fire-killed trees on the Great Tillamook burn in Oregon in 1933 (Furniss 1937b). Figure 6. --Larvae and gnZZeries of '~3ze f Zatheaded f
Other wood borers. --Several species of ambrosia beet1 es, or pinhole borers, are associated with conifers predisposed by wildfire to beetle attack. Usually restricted to the sapwood, ambrosia beetles begin the process of wood deterioration (Beal and others 1935) through a combination of their galleries and an associated staining fungus (Kimmey and Furniss 1943); these beetles usually do not make wood completely useless (Furniss 1937b). In Colorado, some Engelmann spruce trees killed by the Engelmann spruce beetle showed evidence of ambrosia beetles, but most of the defect caused by these borers in the sapwood was removed with the slabs when the trees were cut into 1 umber (Miel ke 1950). Wood wasps (Siricidae) are also attracted to fire-killed trees, and may often severely damage the cuter heartwood (Wallis and others 1971) (fig. 7). Siricids, along with roundheaded borers, produce larger holes than ambrosia beetles and are usually of more economic concern, even though their populations in the same tree may be lower. If lumber cut from fire-salvaged trees is not kiln-dried, siricids may emerge after the product is in use (Wall is and others 1971). Other insect species.--Besides the more abundant and economically important bark beetles and wood borers, wildfires in coniferous forests attract a variety of other insects, some predatory, some scavengers (Boag and Evans 1967). Many species are attracted to fires by smoke and heat (Evans 1971); among those reported to be associated with fire are three species of Empidid or Platypezid smokeflies--Hermope- *, and Microsania occidental is (Komarek 1969) and M. imperfectus (Snoddy and Tippins 1968). RESIDUES CREATED BY WINDTHROW Windthrow represents a natural and often catastrophic event that, like wildfire, predisposes coniferous forests to insects and disease by creating tremendous amounts of residues. Throughout the West, as well as el sewhere on the Continent, several species of bark beetles and wood borers are known to degrade downed timber resulting from windthrow, and to threaten standing timber after breeding in the downed residues (Wickman 1965). All known major outbreaks of the Douglas-fir beetle in western Oregon and Washington have been triggered by severe forest disturbances, particularly by residues created from extensive blowdown during storms (Wright and Harvey 1967). During the winters of 1949-1 950 and 1950-1 951, wind storms blew down some 10 mi 11 ion board feet of sawtimber and in the following 3 years another 3 billion board feet of standing timber were killed by beetles (Wright and Lauderbach 1958). A windstorm of hurricane force struck the Pacific Northwest on October 12, 1962, and in northern California alone blew down nearly a bill ion board feet of coniferous timber. Wickman (1965) caught, reared, or trapped at least 46 different species of insects degrading wood of those windblown trees. Species of Melanophila were the most numerous wood borers, and several species of &and Dendroctonus were the most common bark beetles. As a result of that storm, Dougl3s-fir beetles attacked injured trees (those felled, broken off or leaning), and then spilled over into live surrounding trees (Johnson and Be1 1 uschi 1969). Windthrown ponderosa pine-as we1 1 as logging residue--is a7 so predisposed to attacks by the western pine beetle (Mitchell and Sartwell 1974). Usually there is 1 i ttle population increase in such residues (Patterson 1927; Beal 1935), however, and over the course of many years residue-associated beetle damage has been rather minor.
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ENVIRONMENTAL CONSEQUENCES OF TIMBE
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USDA Forest Service General Technic
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REW 'ORD One of the pressing proble
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BIOLOGICAL IMPLICATIONS Biological
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By the mid-1 960's the apparently u
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Scientist, audience: "The Three-Mil
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I believe that the future of forest
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enewable resource that can be proce
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productive capacity. Recent legisla
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Much of the research investigating
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Six sale area blocks were logged (f
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Understory Removal Figure 4.--Eccpe
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Gentle slopes and easy access to cu
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Each of the four cutting units was
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A concerted effort was made to coor
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oriented toward the basic response
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WOODY MATERIAL IN NORTHERN ROCKY MO
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STANDING, 3 " 1 DIA. ? (GREEN & DEA
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I 11 Table 1,--Volume of wood by co
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Small material under 3 inch (7.6 cm
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Depending on the cutting and treatm
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MICROENVIRONMENTAL RESPONSE TO HARV
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I Where: Cs is the concentration at
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Figure I.-- Residue treatments used
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HARVESTING AND RESIDUE INFLUENCES R
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I 1 1 1 1 1 1 1 1 1 1 1 JAN. FEB. M
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Sensible heat flux, evaporative flu
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SOLO, 1977- 78 UNCUT CUT Mean max.
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It is logical that if surface tempe
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0 - 5 - 10 - 15 20 - Chips - Clrare
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Brown, crumbly, decayed material (B
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At Coram, steep harvested slopes ha
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surface of unburned and hard-burned
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canopy. Light then indirectly cause
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Moisture Moisture is frequently ide
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Net radiation--the measure of the a
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Table 5.--Possible solutions for po
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Cochran, P. H. 1969. Thermal proper
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Lommen, P. W., Cr, R. Schwintzer, C
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Stark, Nel 1 ie. 1980. The impacts
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and water use during the growing se
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The main study site (fig. 1) occupi
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The four residues util i zation tre
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RAIN The major precipitation instru
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Figure 5.--SG,L. wuwr zc)as measure
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S treamf 1 ow A &foot H-flume was i
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Figure 7.--Snow water equivalent on
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Soi 1 Water Water present in the so
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SILVICULTURE TREATMENTS Si 1 vicul
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As indicated in figure 11 and appen
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The most consistent difference in w
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MEAN CONSUMPTIVE USE -ALL YEARS (19
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Lowest daily water use occurred at
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6. Water use began in Apri 1 , acce
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Klages, M. G., R. C. McConnell , an
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APPENDICES Appendix A.--Hydrograph
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Appendix C.--Rainfall reaching the
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Appendix E.--Water use during the g
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One of the most efficient methods o
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Control Chipped Picked up Broadcast
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Control Chapped Picked up Broadcast
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Control Chipped Picked up Broadcast
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' Control Chipped and Picked up and
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One of the least detrimental residu
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INTRODUCTION In the early days, the
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%pire 1.--Measured concentrations (
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The silvicul tural treatments appli
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I 2 Table 2. --Percent of total qua
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Unmerchantable TOP Shrubs Lower Cro
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create a nitrogen deficiency during
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Steele, R. W. 1975. Understory burn
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National Forest. This paper summari
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UNIT 2 CONVENT IONALLY Du Bois, Wyo
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Trees Figures 3 and 4 show the oven
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Weights of typical trees on these t
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pi1 ed-burned treatments were lowes
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MINERAL SOIL Total nutrient content
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Tab1 e 5. --Concentrations of avai
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Table 6. --Average concentrations o
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sites. Burning then transformed the
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RESIDUE DECAY PROCESSES AND ASSOCIA
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The hardwood or angiospermous timbe
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Woody Substrates Gyrnnospermsr (Int
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Natural inoculation of residues may
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On the Coram site, brown-rot fungi
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Table 2.--Probability (P) of encoun
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Recognizable brown-cubical decayed
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I Newly formed, small-dimension res
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Highley, T. L. 1976. Hemicellulases
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MICROBIAL PROCESSES ASSOCIATED WITH
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Nonsymbiotic N Fixation Free-1 ivin
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In a previous paper (Jurgensen and
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Symbiotic N Fixation The establishm
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carbon and NHq (Ahlgren, 1974). Mic
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Prescribed f~re CLEARCUT - BURN CLE
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Hungerford, R.9. 1980. Microenvi ro
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ECOLOGY OF ECTOMYCORRHIZAE IN NORTH
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Coram - Subalpine fir Site (ABLAICL
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Figure 2.-- ReZative yield capabiZi
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Figure 5. -- Percentage of soCZ-woo
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Figure 9,-- Percentage of toid ectm
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Effect of Soil Components on Ectomy
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Effect of Organic Matter Quantity o
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PARTIAL CUT Effect of Harvesting on
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Figure 20. -- Average nwnbers of ac
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Gijbl , F. 1967. Mykorrhizauntersuc
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BIOLOGICAL IMPLICATIONS Initial cha
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FOREST SOIL BIOLOGY The act of incr
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the soil will retain its ability to
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Since the best quantity of organic
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We can also visualize situations on
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Harvey, A. E., M. F. Jurgensen, and
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INTRODUCTION Vegetation integrates
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Because of cool, wet weather in 197
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We determined volume and cover usin
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The stems were divided at 4 mm (0.1
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RESIDUES TREATMENT EFFECTS As descr
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Overall shrub recovery rates do not
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Rose behaved similar to ninebark (f
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Meanwhile, small shrub cover change
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Biomass Regression analyses relatin
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SILVICULTURE TREATMENT EFFECTS Harv
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RESIDUES TREATMENT EFFECTS Major sh
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Lesser vegetation--herbs and small
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Leaf and stem components of the shr
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APPENDIX I Trees and shrubs found o
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INTRODUCTION Many western larch (La
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Figure 2. --Lower cutting blocks on
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each permanent point) of the clearc
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Figure 3.--Marking gemination and c
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were not significant. These treatme
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Tab1 e 2. --Fi 11 ed conifer seed (
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Dispersal of sound seed on the uppe
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Table $.--Number of 3- and 5-year-o
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Table 6. --Germination of conifers
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Survival, growth and form will be m
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Es tab1 ishment and Initial Develop
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to the amount of residue originally
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As the following tabulation shows,
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Soot-Seeded Broadcast burning, and
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Natural Regeneration Natural regene
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.4s shown in fig. 5, vegetation was
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Interestingly, the high phenol leve
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DeByle, Norbert V. 1980. Harvesting
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EFFECT OF SILVICULTURAL PRACTICES,
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mountain pine beetle. These two for
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GROUP SELECTION 22 CONTROL SHELTERW
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FIELD AND LABORATORY PROCEDURES To
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I Many of the groups were trapped v
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shelterwood with residues, were sig
Page 301 and 302: HARVESTING AND RESIDUE MANAGEMENT T
Page 307 and 308: indicate a treatment effect of burn
Page 309 and 310: Undisturbed Forest .". . . . Clearc
Page 311 and 312: 1 equally abundant in all treatment
Page 313 and 314: I not seem to be directly caused by
Page 315 and 316: the undisturbed forests. Some resea
Page 317 and 318: Huhta, V., E. Karppinen, M. Nurmine
Page 319 and 320: POPULATIONS OF SOME FOREST LITTER,
Page 321 and 322: STUDY DESIGN The study area is loca
Page 323 and 324: GROUP SELECTION 22 CONTROL '... . .
Page 325 and 326: Sampling in 1977 was restricted to
Page 327 and 328: Table 1 .--Mean total mesofauna pop
Page 329 and 330: 0 - (JUNE) from S helterwood (AUGUS
Page 331 and 332: SHELTERWOOD--RESIDUE BURNED The eff
Page 333 and 334: In the first season or two followin
Page 335 and 336: LITERATURE CITED Ahlgren, I. F. 197
Page 337 and 338: A REVIEW OF SOME INTERACTIONS BETWE
Page 339 and 340: and Keen 1960; Felix and others 197
Page 341 and 342: Pissodes strobi Peck, In a thinned
Page 343 and 344: foresters feel as though "It's 1 i
Page 345 and 346: The large aerial spray programs aga
Page 347 and 348: Weakened living trees, or trees kil
Page 349 and 350: Most western pine beetle attacks in
Page 351: The galleries, or mines, of wood bo
Page 355 and 356: Deterioration of spruce (Picea a. )
Page 357 and 358: The eruption of Mount Saint Helens
Page 359 and 360: Bark and engraver beetles infesting
Page 361 and 362: In the northern Rockies, one conife
Page 363 and 364: management, and not the beetle." If
Page 365 and 366: investigation regardin controlled b
Page 367 and 368: WOOD BORERS Mitchell and Martin (In
Page 369 and 370: Still, by 1938, there was a feeling
Page 371 and 372: I Different i nvesti gators have us
Page 373 and 374: if the forest floor is completely c
Page 375 and 376: Carabids as biological control agen
Page 377 and 378: een placed for protection from rode
Page 379 and 380: with every tree in the stand contin
Page 381 and 382: Spiders as predators.--Studies have
Page 383 and 384: At least two studies have been made
Page 385 and 386: In the long-leaf pine forests in th
Page 387 and 388: RESIDUES, FIRES, INSECTS, AND FORES
Page 389 and 390: In the spruce-fir stands infested w
Page 391 and 392: Recently, Senator John Me1 cher (Mo
Page 393 and 394: plans. Since fire, too, is a decomp
Page 395 and 396: CHANGING FOREST INSECT PROBLEMS Ins
Page 397 and 398: Basham, J. T. 1957. The deteriorati
Page 399 and 400: Cole, D. M. 1978. Feasibility of si
Page 401 and 402: Fellin, D. G. and P. C. Johnson. 19
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Graham, S. A. 1922. Some entomoloqi
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Huhta, V., M. Nurminen and A. Valpa
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Lawrence, W. H. and 3. H. Rediske.
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Miller, J. M. and J. E. Patterson.
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Rice, L * A. 1932. The effect of fi
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Stoddard, H. L. Sr. 1963. Bird habi
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Weaver, Harol d. 1951. Fire as an e
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RESOURCE MANAGEMENT IMPLICATIONS Th
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INFLUENCE OF HARVESTING AND RESIDUE
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F
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NEAR COMPLETE Figure 2.--Byrmnts f
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Figure 3.-- Bymrmrs fireZine intens
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F5qure 4. --Byramrs fireZ-Cne inten
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1. Nomographs of Rate of Spread, Fi
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3. Consider other fire-related fact
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Puckett, John V., Cameron M. Johnst
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Important and rapid progress has be
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tlarvested Areas What happened when
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UNCUT SHELTERWOOD Protect understor
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Initially the areas are rated low.
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The primary reason for this discrep
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f n contrast to displaced bears, so
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Quantitative data on grizzly behavi
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Asherin, Duane A. 1976. Changes in
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Leege, Thomas A. 1969. Burning sera
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Reynolds, Hudson G. 1969. Aspen gro
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INFLUENCES OF HARVESTING AND RESIDU
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CLEARCUT LOGGING & COMPLETE BURN DE
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C LEARCUT LOGGING & BROADCAST BURN
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EFFECTS OF SMALL MAMMALS ON FOREST
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I I Habitat Manipulation Small mamm
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Fala, Robert A. 1975. Effects of pr
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Tevis, Lloyd Jr. l956b. Pocket goph
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forests are cavity nesters. They ar
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GROUP SELECTION: A1 1 merchantab le
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TABLE 3. Pre- and post-logging volu
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Censuses Censuses for all species (
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Fbgure 6.-- Lightn
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Pileated Woodpeckers on the CEF nes
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TABLE 7. Percent of sampling time t
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I Observed woodpecker feeding Down,
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Table 9 shows feeding time in the l
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L c - 10- ul 2 0 8- 6 - 4 2 - - 0 .
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Figure 15. -- A Hairy Woodpecker ne
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~
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oth offered advice on study site se
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Hairy Woodpecker Downy Woodpecker B
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THE SITUATION Forest land managers
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In the descriptive approach, howeve
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ORGANIZING INFORMATION One differen
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As has already been stated, the for
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Figure 7.--Matrix of interactions b
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understory biomass density, X2 repr
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or, by rearrangement, AX. I afi Ax,
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DISCUSSION It is important to note
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LITERATURE CITED Billings, W. D. 19
Page 515 and 516:
Richard T. Wick Burlington Northern
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themselves, the traditional industr
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highways. Of course, all roads are
Page 521 and 522:
Darrel L. Kenops District Ranger US
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In the future we will see our const
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Lyon, L. Jack, Project Leader, Ecol
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The Intermountain Station, headquar
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