ENVIRONMENTAL CONSEQUENCES in rocky mountain coniferous ...

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The primary reason for larch selection and apparent avoidance of Douglas-fir is related to decay characteristics of the two species. After tree death, Douglas-fir sapwood decays rapidly along with the heartwood (Wright and Harvey 1967). A protec- tive "shell" of sapwood is missing in snags of that species. In western larch, the sapwood decays very slowly, perpetuating a shell around decaying heartwood for many years. Cavities in Douglas-fir deteriorate more rapidly and are more susceptible to predation (a predator is able to gain access to the nest chamber by enlarging the entrance, which is in decaying wood). No live Douglas-fir were used as nest sites, but 1 ive larch were used more of ten than were larch snags. Nest cavities in 1 ive larch were usually excavated through living wood tissue into decaying heartwood rather than in a dead top or a dead branch. With one exception (a Black-backed Three -toed Woodpecker nest), Douglas-fir snags were used only by the "weak" excavators (flickers, chickadees, and nuthatches). They seem to require very soft wood for successful completion of cavities. In instances where the weak excavators nested in trees species such as western larch, they typically used cavities abandoned by stronger excavators. Mountain Chickadees and Red-breasted Nuthatches usually nested in old nest holes of Yellow-bellied Sapsuckers (woodpeckers nearly always excavate a new nest cavity each spring). In western larch, selection related to tree condition was directed toward broken tops (fig. 5). Although there were nearly equal numbers of intact-top and broken-top larch snags (and mean dbh of both groups exceeded 15 in (38 crn)) , there were about 5 times more nests in broken-top snaqs. This selection process probably involves decay presence. A broklen top provides an avenue of entry for decay orqanisms (Hepting 1971 ) . Top breakage in western larch commonly resul ts from 1 iqhtninq (fig. 6), snow or ice accumulation, or windstorm. Broken tops also may result from breakage at a point weakened by pre-existinp decay or the presence of a nest cavity (Conner et a1 . 1975). 1 1 Relative density BE Nest trees In = 183) . . .G I top top 1 I fop dead top live top I SNAGS L l VE TREES TREE CONDITION Figure 5. -- Dee corzdikion preferences, western larch nest trees, all hole-nesting bird species cornb
Fbgure 6.-- Lightn
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ENVIRONMENTAL CONSEQUENCES OF TIMBE
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USDA Forest Service General Technic
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REW 'ORD One of the pressing proble
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BIOLOGICAL IMPLICATIONS Biological
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By the mid-1 960's the apparently u
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Scientist, audience: "The Three-Mil
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I believe that the future of forest
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enewable resource that can be proce
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productive capacity. Recent legisla
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Much of the research investigating
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Six sale area blocks were logged (f
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Understory Removal Figure 4.--Eccpe
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Gentle slopes and easy access to cu
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Each of the four cutting units was
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A concerted effort was made to coor
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oriented toward the basic response
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WOODY MATERIAL IN NORTHERN ROCKY MO
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STANDING, 3 " 1 DIA. ? (GREEN & DEA
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I 11 Table 1,--Volume of wood by co
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Small material under 3 inch (7.6 cm
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Depending on the cutting and treatm
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MICROENVIRONMENTAL RESPONSE TO HARV
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I Where: Cs is the concentration at
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Figure I.-- Residue treatments used
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HARVESTING AND RESIDUE INFLUENCES R
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I 1 1 1 1 1 1 1 1 1 1 1 JAN. FEB. M
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Sensible heat flux, evaporative flu
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SOLO, 1977- 78 UNCUT CUT Mean max.
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It is logical that if surface tempe
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0 - 5 - 10 - 15 20 - Chips - Clrare
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Brown, crumbly, decayed material (B
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At Coram, steep harvested slopes ha
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surface of unburned and hard-burned
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canopy. Light then indirectly cause
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Moisture Moisture is frequently ide
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Net radiation--the measure of the a
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Table 5.--Possible solutions for po
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Cochran, P. H. 1969. Thermal proper
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Lommen, P. W., Cr, R. Schwintzer, C
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Stark, Nel 1 ie. 1980. The impacts
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and water use during the growing se
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The main study site (fig. 1) occupi
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The four residues util i zation tre
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RAIN The major precipitation instru
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Figure 5.--SG,L. wuwr zc)as measure
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S treamf 1 ow A &foot H-flume was i
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Figure 7.--Snow water equivalent on
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Soi 1 Water Water present in the so
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SILVICULTURE TREATMENTS Si 1 vicul
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As indicated in figure 11 and appen
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The most consistent difference in w
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MEAN CONSUMPTIVE USE -ALL YEARS (19
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Lowest daily water use occurred at
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6. Water use began in Apri 1 , acce
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Klages, M. G., R. C. McConnell , an
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APPENDICES Appendix A.--Hydrograph
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Appendix C.--Rainfall reaching the
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Appendix E.--Water use during the g
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One of the most efficient methods o
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Control Chipped Picked up Broadcast
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Control Chapped Picked up Broadcast
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Control Chipped Picked up Broadcast
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' Control Chipped and Picked up and
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One of the least detrimental residu
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INTRODUCTION In the early days, the
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%pire 1.--Measured concentrations (
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The silvicul tural treatments appli
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I 2 Table 2. --Percent of total qua
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Unmerchantable TOP Shrubs Lower Cro
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create a nitrogen deficiency during
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Steele, R. W. 1975. Understory burn
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National Forest. This paper summari
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UNIT 2 CONVENT IONALLY Du Bois, Wyo
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Trees Figures 3 and 4 show the oven
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Weights of typical trees on these t
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pi1 ed-burned treatments were lowes
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MINERAL SOIL Total nutrient content
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Tab1 e 5. --Concentrations of avai
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Table 6. --Average concentrations o
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sites. Burning then transformed the
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RESIDUE DECAY PROCESSES AND ASSOCIA
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The hardwood or angiospermous timbe
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Woody Substrates Gyrnnospermsr (Int
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Natural inoculation of residues may
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On the Coram site, brown-rot fungi
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Table 2.--Probability (P) of encoun
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Recognizable brown-cubical decayed
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I Newly formed, small-dimension res
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Highley, T. L. 1976. Hemicellulases
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MICROBIAL PROCESSES ASSOCIATED WITH
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Nonsymbiotic N Fixation Free-1 ivin
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In a previous paper (Jurgensen and
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Symbiotic N Fixation The establishm
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carbon and NHq (Ahlgren, 1974). Mic
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Prescribed f~re CLEARCUT - BURN CLE
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Hungerford, R.9. 1980. Microenvi ro
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ECOLOGY OF ECTOMYCORRHIZAE IN NORTH
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Coram - Subalpine fir Site (ABLAICL
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Figure 2.-- ReZative yield capabiZi
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Figure 5. -- Percentage of soCZ-woo
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Figure 9,-- Percentage of toid ectm
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Effect of Soil Components on Ectomy
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Effect of Organic Matter Quantity o
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PARTIAL CUT Effect of Harvesting on
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Figure 20. -- Average nwnbers of ac
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Gijbl , F. 1967. Mykorrhizauntersuc
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BIOLOGICAL IMPLICATIONS Initial cha
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FOREST SOIL BIOLOGY The act of incr
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the soil will retain its ability to
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Since the best quantity of organic
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We can also visualize situations on
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Harvey, A. E., M. F. Jurgensen, and
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INTRODUCTION Vegetation integrates
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Because of cool, wet weather in 197
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We determined volume and cover usin
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The stems were divided at 4 mm (0.1
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RESIDUES TREATMENT EFFECTS As descr
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Overall shrub recovery rates do not
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Rose behaved similar to ninebark (f
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Meanwhile, small shrub cover change
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Biomass Regression analyses relatin
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SILVICULTURE TREATMENT EFFECTS Harv
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RESIDUES TREATMENT EFFECTS Major sh
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Lesser vegetation--herbs and small
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Leaf and stem components of the shr
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APPENDIX I Trees and shrubs found o
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INTRODUCTION Many western larch (La
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Figure 2. --Lower cutting blocks on
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each permanent point) of the clearc
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Figure 3.--Marking gemination and c
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were not significant. These treatme
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Tab1 e 2. --Fi 11 ed conifer seed (
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Dispersal of sound seed on the uppe
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Table $.--Number of 3- and 5-year-o
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Table 6. --Germination of conifers
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Survival, growth and form will be m
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Es tab1 ishment and Initial Develop
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to the amount of residue originally
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As the following tabulation shows,
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Soot-Seeded Broadcast burning, and
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Natural Regeneration Natural regene
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.4s shown in fig. 5, vegetation was
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Interestingly, the high phenol leve
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DeByle, Norbert V. 1980. Harvesting
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EFFECT OF SILVICULTURAL PRACTICES,
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mountain pine beetle. These two for
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GROUP SELECTION 22 CONTROL SHELTERW
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FIELD AND LABORATORY PROCEDURES To
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I Many of the groups were trapped v
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shelterwood with residues, were sig
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HARVESTING AND RESIDUE MANAGEMENT T
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indicate a treatment effect of burn
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Undisturbed Forest .". . . . Clearc
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1 equally abundant in all treatment
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I not seem to be directly caused by
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the undisturbed forests. Some resea
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Huhta, V., E. Karppinen, M. Nurmine
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POPULATIONS OF SOME FOREST LITTER,
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STUDY DESIGN The study area is loca
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GROUP SELECTION 22 CONTROL '... . .
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Sampling in 1977 was restricted to
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Table 1 .--Mean total mesofauna pop
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0 - (JUNE) from S helterwood (AUGUS
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SHELTERWOOD--RESIDUE BURNED The eff
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In the first season or two followin
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LITERATURE CITED Ahlgren, I. F. 197
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A REVIEW OF SOME INTERACTIONS BETWE
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and Keen 1960; Felix and others 197
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Pissodes strobi Peck, In a thinned
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foresters feel as though "It's 1 i
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The large aerial spray programs aga
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Weakened living trees, or trees kil
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Most western pine beetle attacks in
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The galleries, or mines, of wood bo
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Other wood borers. --Several specie
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Deterioration of spruce (Picea a. )
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The eruption of Mount Saint Helens
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Bark and engraver beetles infesting
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In the northern Rockies, one conife
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management, and not the beetle." If
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investigation regardin controlled b
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WOOD BORERS Mitchell and Martin (In
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Still, by 1938, there was a feeling
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I Different i nvesti gators have us
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if the forest floor is completely c
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Carabids as biological control agen
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een placed for protection from rode
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with every tree in the stand contin
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Spiders as predators.--Studies have
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At least two studies have been made
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In the long-leaf pine forests in th
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RESIDUES, FIRES, INSECTS, AND FORES
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In the spruce-fir stands infested w
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Recently, Senator John Me1 cher (Mo
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plans. Since fire, too, is a decomp
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CHANGING FOREST INSECT PROBLEMS Ins
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Basham, J. T. 1957. The deteriorati
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Cole, D. M. 1978. Feasibility of si
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Fellin, D. G. and P. C. Johnson. 19
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Graham, S. A. 1922. Some entomoloqi
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Huhta, V., M. Nurminen and A. Valpa
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Lawrence, W. H. and 3. H. Rediske.
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Miller, J. M. and J. E. Patterson.
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Rice, L * A. 1932. The effect of fi
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Stoddard, H. L. Sr. 1963. Bird habi
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Weaver, Harol d. 1951. Fire as an e
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RESOURCE MANAGEMENT IMPLICATIONS Th
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INFLUENCE OF HARVESTING AND RESIDUE
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F
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NEAR COMPLETE Figure 2.--Byrmnts f
Page 425 and 426: Figure 3.-- Bymrmrs fireZine intens
Page 427 and 428: F5qure 4. --Byramrs fireZ-Cne inten
Page 429 and 430: 1. Nomographs of Rate of Spread, Fi
Page 431 and 432: 3. Consider other fire-related fact
Page 433 and 434: Puckett, John V., Cameron M. Johnst
Page 435 and 436: Important and rapid progress has be
Page 437 and 438: tlarvested Areas What happened when
Page 439 and 440: UNCUT SHELTERWOOD Protect understor
Page 441 and 442: Initially the areas are rated low.
Page 443 and 444: The primary reason for this discrep
Page 445 and 446: f n contrast to displaced bears, so
Page 447 and 448: Quantitative data on grizzly behavi
Page 449 and 450: Asherin, Duane A. 1976. Changes in
Page 451 and 452: Leege, Thomas A. 1969. Burning sera
Page 453 and 454: Reynolds, Hudson G. 1969. Aspen gro
Page 455 and 456: INFLUENCES OF HARVESTING AND RESIDU
Page 457 and 458: CLEARCUT LOGGING & COMPLETE BURN DE
Page 459 and 460: C LEARCUT LOGGING & BROADCAST BURN
Page 461 and 462: EFFECTS OF SMALL MAMMALS ON FOREST
Page 463 and 464: I I Habitat Manipulation Small mamm
Page 465 and 466: Fala, Robert A. 1975. Effects of pr
Page 467 and 468: Tevis, Lloyd Jr. l956b. Pocket goph
Page 469 and 470: forests are cavity nesters. They ar
Page 471 and 472: GROUP SELECTION: A1 1 merchantab le
Page 473 and 474: TABLE 3. Pre- and post-logging volu
Page 475: Censuses Censuses for all species (
Page 479 and 480: Pileated Woodpeckers on the CEF nes
Page 481 and 482: TABLE 7. Percent of sampling time t
Page 483 and 484: I Observed woodpecker feeding Down,
Page 485 and 486: Table 9 shows feeding time in the l
Page 487 and 488: L c - 10- ul 2 0 8- 6 - 4 2 - - 0 .
Page 489 and 490: Figure 15. -- A Hairy Woodpecker ne
Page 491 and 492: ~
Page 493 and 494: oth offered advice on study site se
Page 495 and 496: Hairy Woodpecker Downy Woodpecker B
Page 497 and 498: THE SITUATION Forest land managers
Page 499 and 500: In the descriptive approach, howeve
Page 501 and 502: ORGANIZING INFORMATION One differen
Page 503 and 504: As has already been stated, the for
Page 505 and 506: Figure 7.--Matrix of interactions b
Page 507 and 508: understory biomass density, X2 repr
Page 509 and 510: or, by rearrangement, AX. I afi Ax,
Page 511 and 512: DISCUSSION It is important to note
Page 513 and 514: LITERATURE CITED Billings, W. D. 19
Page 515 and 516: Richard T. Wick Burlington Northern
Page 517 and 518: themselves, the traditional industr
Page 519 and 520: highways. Of course, all roads are
Page 521 and 522: Darrel L. Kenops District Ranger US
Page 523 and 524: In the future we will see our const
Page 525 and 526: Lyon, L. Jack, Project Leader, Ecol
Page 527:
The Intermountain Station, headquar
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