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ENVIRONMENTAL CONSEQUENCES in rocky mountain coniferous ...

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Seed Loss After Dispersal<br />

From the time tree seed reaches the ground <strong>in</strong> the fall until it germ<strong>in</strong>ates the<br />

foll ow<strong>in</strong>g spr<strong>in</strong>g, many factors (chiefly rodents, birds, <strong>in</strong>vertebrates and diseases)<br />

reduce the number of seed potentially capable of germ<strong>in</strong>ation. The number of viable<br />

seeds was estimated from seed traps. Seed loss dur<strong>in</strong>g w<strong>in</strong>ter was estimated from 6-<br />

square-foot (0.5 m2) samples of duff and soil taken from surfaces about 10 feet (3 m)<br />

from selected seed traps. These samples were taken <strong>in</strong> May 1475 at 16 po<strong>in</strong>ts with<strong>in</strong><br />

each cutt<strong>in</strong>g method and with<strong>in</strong> the control (uncut) areas,<br />

The difference between the number of filled seed found <strong>in</strong> the seed traps and<br />

those found at the surface represents an estimate of seed loss.<br />

Seedl<strong>in</strong>g Establishment<br />

Seed1 <strong>in</strong>g establishment is <strong>in</strong>fluenced not only by the number of seeds that<br />

germ<strong>in</strong>ate but by a broad range of biological and phys ical factors affectieg surv<br />

particularly dur<strong>in</strong>g the first year fol low<strong>in</strong>g germ<strong>in</strong>at ion.<br />

Establishment of natural regeneration was studied on permanent circular milacre<br />

plots (43.56 ft2, or 4.05 m2, <strong>in</strong> surface area) with<strong>in</strong> each subblock; the locations of<br />

these plots were chosen to reflect the conditions of the treatment areas <strong>in</strong> which<br />

they were <strong>in</strong>stalled. The center of each plot was located 23 feet (7 m). from each<br />

permanent po<strong>in</strong>t at the lower right corner (look<strong>in</strong>g uphill 1 of the 53.82 ft2 (5 m2)<br />

vegetation quadrants. Measurements were made <strong>in</strong> the summers of 1978 and 1979. In<br />

1979, the sample size was doubled by establish<strong>in</strong>g temporary po<strong>in</strong>ts 50 feet (15.24 m)<br />

downhill from each permanent po<strong>in</strong>t. Seed-seedl<strong>in</strong>g ratios were estimated us<strong>in</strong>g the<br />

seed trap closest to each natural regeneration plot.<br />

Seeded Plots<br />

In order to identify the probable causes of seedl<strong>in</strong>g mortality, plots were<br />

established on each of the seedbed and residue subblocks with<strong>in</strong> all cutt<strong>in</strong>g blocks<br />

(fig. 3). Two and four plots were seeded on each of the burned and unburned sub-<br />

block, respectively. These plots were sown with 200 seeds of each of the follow<strong>in</strong>g<br />

species: western larch, Douglas-fir, Engelmann spruce, subalp<strong>in</strong>e fir, and lodgepol e<br />

p<strong>in</strong>e ( P<strong>in</strong>us contorta Doug1 . ex Loua. ) . A1 1 plots were sown from mid- to late-October<br />

for 5 consecutive years (1974 - 1978), except those that were unburned <strong>in</strong> 1974 on<br />

subunits 1 and 2 of the shelterwood cutt<strong>in</strong>g (block 11), shown <strong>in</strong> fig. 3.<br />

These plots were located and marked as soon as possible after the treatments<br />

were completed--<strong>in</strong> the fa1 7 of 1974 on unburned seedbeds and <strong>in</strong> the fa1 1 of 1975 on<br />

burned seedbeds,<br />

The seeds used <strong>in</strong> this study were collected near the Coram Experimental Forest<br />

<strong>in</strong> the fall of 1974, <strong>in</strong> an area of the same habitat type, elevational zone and<br />

aspect as the study area. Extraction and clean<strong>in</strong>g was done at the Coeur dlA]ene<br />

nursery; chemical repellent coat<strong>in</strong>g was not used, so that field germ<strong>in</strong>ation would be<br />

as similar to that of naturally dispersed seed as possible.

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