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ENVIRONMENTAL CONSEQUENCES in rocky mountain coniferous ...

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By chang<strong>in</strong>g w<strong>in</strong>d patterns, clearcutt<strong>in</strong>g and partial cutt<strong>in</strong>g <strong>in</strong>crease the suscep-<br />

ti bil i t y of a stand to blowdown. Considerable w<strong>in</strong>dthrow occurred around the clearcuts<br />

and with<strong>in</strong> the partial cuts at Coram. Partially-cut sites on the Flathead National<br />

Forest, at least <strong>in</strong> the vic<strong>in</strong>ity of Hungry Horse Reservoir, frequently experience<br />

bl owdown,<br />

Residues left on the site slow air movement near the ground. The amount,<br />

character, and placement of residues alter the w<strong>in</strong>d speed with height. As a result,<br />

evaporation and convective energy exchanges are a1 tered. Decreas<strong>in</strong>g w<strong>in</strong>d movement<br />

decreases m<strong>in</strong>imum and <strong>in</strong>creases maximum temperatures.<br />

POTENTIAL BIOLOGICAL <strong>CONSEQUENCES</strong><br />

Light<br />

The amount, qua1 ity, and duration of 1 ight are significant to biological pro-<br />

cesses, Light affects plant growth through both photosynthesis and the thermal<br />

balance that controls transpiration and other <strong>in</strong>ternal physiological processes.<br />

Light levels also affect the distribution and survival of other organisms,<br />

Photosynthesis depends on a photochemical reaction that requires 7 ight energy to<br />

split water molecules. If other factors of heat and COz are not 1 imit<strong>in</strong>g, photosyn-<br />

thesis <strong>in</strong>creases <strong>in</strong> proportion to 1 ight <strong>in</strong>tensity. Species have different <strong>in</strong>tensity<br />

requirements at which maximum photosynthesis will occur. Foresters have labeled<br />

plants tolerant or <strong>in</strong>tolerant. Certa<strong>in</strong> of the chemical processes <strong>in</strong> photosynthesis<br />

are temperature sensitive or COz sensitive. If either of these factors becomes<br />

critical under adequate 1 ight, photosynthesis will decrease. Any one of these three<br />

factors may become limit<strong>in</strong>g <strong>in</strong> nature. Water is also critical to the photosynthetic<br />

rate which can be limited by dry or cold soils. The viscosity of water is <strong>in</strong>creased<br />

<strong>in</strong> cold soils; this <strong>in</strong>creases the resistance to movement and decreases the rates of<br />

chemical reactions. High 1 ight <strong>in</strong>tensities and high temperatures <strong>in</strong>directly 1 i mi t<br />

photosynthesis by caus<strong>in</strong>g damage to tissues. Excessive supplies of COP cause <strong>in</strong>jury<br />

a1 so.<br />

Photosynthesis and respiration determ<strong>in</strong>e growth rate and production of biomass.<br />

Respiration rates or oxidation of the carbohydrates produced <strong>in</strong> photosynthesis are<br />

controlled by temperature asd oqgen levels. Plants with photorespiration exhibit<br />

optimum respiration fro! 10 C-25 C. Plants without photorespiration exhibit optimum<br />

respiration at about 35 C. Increases <strong>in</strong> temperature at certa<strong>in</strong> levels <strong>in</strong>crease<br />

respiration and decrease net photosynthesis. Lee and Sypol t (1974) found that basal<br />

area growth of a forest <strong>in</strong> West Virg<strong>in</strong>ia was reduced because high canopy temperatures<br />

reduced dry matter production. These high canopy temperatures resulted from high net<br />

radiation loads and cooler soi 1 temperatures, The cooler soil temperatures decreased<br />

water absorption to the po<strong>in</strong>t where canopy transpiration exceeded the rate of water<br />

absorption. Lee and Sypolt (1974) suggested that high net radiation levels (greater<br />

than 0.7 cal/cm2/m<strong>in</strong>) may directly reduce net assimilation.<br />

Light, as <strong>in</strong>dicated <strong>in</strong> the above discussion, can <strong>in</strong>crease the temperature of a<br />

plant leaf. This <strong>in</strong>crease <strong>in</strong> temperature <strong>in</strong>creases the difference between the vapor<br />

pressures of air and of the <strong>in</strong>ternal cavities of the leaf, thereby, <strong>in</strong>creas<strong>in</strong>g<br />

transpiration. This is what took place <strong>in</strong> the above example. Light also causes the<br />

stomates to open, allow<strong>in</strong>g transpiration. Transpiration effectively cools the

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