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Carbaryl, Carbofuran, and Methomyl - National Marine Fisheries ...

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ground water <strong>and</strong> degrade water quality in the freshwater, estuarine, <strong>and</strong> coastal<br />

ecosystems throughout the Pacific Northwest.<br />

Coho Salmon<br />

Description of the Species<br />

Coho salmon occur naturally in most major river basins around the North Pacific Ocean<br />

from central California to northern Japan (Laufle, Pauley et al. 1986). We discuss the<br />

distribution, life history diversity, status, <strong>and</strong> critical habitat of the four endangered <strong>and</strong><br />

threatened coho species separately.<br />

After entering the ocean, immature coho salmon initially remain in nearshore waters<br />

close to the parent stream. Most coho salmon adults are three-year-olds, having spent<br />

approximately 18 months rearing in freshwater <strong>and</strong> 18 months in salt water. Most coho<br />

salmon enter rivers between September <strong>and</strong> February. However, entry is influenced by<br />

discharge <strong>and</strong> other factors. In many systems, coho salmon <strong>and</strong> other Pacific salmon are<br />

unable to enter the rivers until sufficiently strong flows open passages <strong>and</strong> provide<br />

sufficient depth. Wild female coho salmon return to spawn almost exclusively at age<br />

three. Coho salmon spawn from November to January, <strong>and</strong> occasionally into February<br />

<strong>and</strong> March. Spawning occurs in a few third-order streams. Most spawning activity<br />

occurs in fourth- <strong>and</strong> fifth-order streams. Spawning generally occurs in tributaries with<br />

gradients of 3% or less.<br />

Eggs incubate for about 35 to 50 days, <strong>and</strong> start emerging from the gravel within two to<br />

three weeks after hatching. Following emergence, fry move to shallow areas near the<br />

stream banks. As fry grow, they disperse upstream <strong>and</strong> downstream to establish <strong>and</strong><br />

defend territories. Juvenile rearing usually occurs in tributaries with gradients of 3% or<br />

less, although they may move to streams with gradients of 4 to 5%. Juvenile coho<br />

salmon are often found in small streams less than five ft wide, <strong>and</strong> may migrate<br />

considerable distances to rear in lakes <strong>and</strong> off-channel ponds. During the summer, fry<br />

prefer pools featuring adequate cover such as large woody debris, undercut banks, <strong>and</strong><br />

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