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Carbaryl, Carbofuran, and Methomyl - National Marine Fisheries ...

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may affect steelhead migration patterns. For example, some populations of coho <strong>and</strong><br />

Chinook salmon, at least historically, remained within Puget Sound <strong>and</strong> did not migrate<br />

to the Pacific Ocean. Even when Puget Sound steelhead migrate to the high seas, they<br />

may spend considerable time as juveniles or adults in the protected marine environment<br />

of Puget Sound. This is a feature not readily accessible to steelhead from other areas of<br />

the Pacific Northwest. The species is primarily composed of winter steelhead but<br />

includes several stocks of summer steelhead, usually in subbasins of large river systems<br />

<strong>and</strong> above seasonal hydrologic barriers.<br />

Life History<br />

Life history attributes of Puget Sound steelhead (migration <strong>and</strong> spawn timing, smolt age,<br />

ocean age, <strong>and</strong> total age at first spawning) appear similar to those of other west coast<br />

steelhead. Ocean age for Puget Sound summer steelhead varies among populations.<br />

Status <strong>and</strong> Trends<br />

Puget Sound steelhead were listed as a threatened species on May 11, 2007 (72 FR<br />

26722). Run size for this DPS, was calculated in the early 1980s at about 100,000<br />

winter-run fish <strong>and</strong> 20,000 summer-run fish. It is unclear what portion were hatchery<br />

fish. However, a combined estimate with coastal steelhead suggested that roughly 70%<br />

of steelhead in ocean runs were of hatchery origin. The percentage in escapement to<br />

spawning grounds would be substantially lower due to differential harvest <strong>and</strong> hatchery<br />

rack returns. By the 1990s, total run size for four major stocks exceeded 45,000, roughly<br />

half of which was natural escapement.<br />

Nehlsen et al. (1991) identified nine Puget Sound steelhead stocks at some degree of risk<br />

or concern. The WDFW et al. (1993) estimated that 31 of 53 stocks were of native origin<br />

<strong>and</strong> predominantly natural production. The WDFW assessment of the status of these 31<br />

stocks was 11 healthy, three depressed, one critical, <strong>and</strong> 16 of unknown status. Their<br />

assessment of the status of the remaining (not native/natural) stocks was three healthy, 11<br />

depressed, <strong>and</strong> eight of unknown status.<br />

165

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