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Carbaryl, Carbofuran, and Methomyl - National Marine Fisheries ...

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channel or from the shoreline of a st<strong>and</strong>ing body of water. Designated critical habitat<br />

areas include the Columbia River from a straight line connecting the west end of the<br />

Clatsop jetty (Oregon side) <strong>and</strong> the west end of the Peacock jetty (Washington side), all<br />

river reaches from the estuary upstream to the confluence of the SR, <strong>and</strong> all SR reaches<br />

upstream to the confluence of the Salmon River; all Salmon River reaches to Alturas<br />

Lake Creek; Stanley, Redfish, yellow Belly, Pettit, <strong>and</strong> Alturas Lakes (including their<br />

inlet <strong>and</strong> outlet creeks); Alturas Lake Creek <strong>and</strong> that portion of Valley Creek between<br />

Stanley Lake Creek; <strong>and</strong> the Salmon River. Limiting factors identified for SR sockeye<br />

include: (1) Reduced tributary stream flow, (2) impaired tributary passage <strong>and</strong> blocks to<br />

migration, (3) degraded water quality; <strong>and</strong> (4) mainstem Columbia River hydropower<br />

system mortality.<br />

Steelhead<br />

Description of the Species<br />

Steelhead are native to Pacific Coast streams extending from Alaska south to<br />

northwestern Mexico (Moyle 1976; Gustafson, Wainwright et al. 1997; Good, Waples et<br />

al. 2005). We discuss the distribution, life history diversity, status, <strong>and</strong> critical habitat of<br />

the 11 endangered <strong>and</strong> threatened steelhead species separately.<br />

Steelhead can be divided into two basic run-types: the stream-maturing type, or summer<br />

steelhead <strong>and</strong> the ocean-maturing type, or winter steelhead. The stream-maturing type or<br />

summer steelhead enters fresh water in a sexually immature condition. It requires several<br />

months in freshwater to mature <strong>and</strong> spawn. The ocean-maturing type or winter steelhead<br />

enters freshwater with well-developed gonads <strong>and</strong> spawns shortly after river entry.<br />

Variations in migration timing exist between populations. Some river basins have both<br />

summer <strong>and</strong> winter steelhead, while others only have one run-type.<br />

Summer steelhead enter freshwater between May <strong>and</strong> October in the Pacific Northwest<br />

(Nickelsen, Nicholas et al. 1992; Busby, Wainwright et al. 1996). They require cool,<br />

deep holding pools during summer <strong>and</strong> fall, prior to spawning (Nickelsen, Nicholas et al.<br />

1992). They migrate inl<strong>and</strong> toward spawning areas, overwinter in the larger rivers,<br />

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