Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
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uses a mutant form <strong>of</strong> lysozyme to digest the bacteria. This<br />
adaptation has evolved separately in cows, langur monkeys,<br />
and hoatzin birds.<br />
In many cases, convergent evolution does not require a total<br />
reinvention <strong>of</strong> an adaptation.<br />
• Limonene has evolved separately in several lineages <strong>of</strong><br />
plants (see above), but most flowering plants have terpene<br />
synthases, the enzymes that produce the immediate chemical<br />
precursors <strong>of</strong> limonene.<br />
• Ornithologist N. I. Mundy explains that the evolution <strong>of</strong><br />
similar color patterns in separate bird lineages has resulted<br />
from the modification <strong>of</strong> an underlying genetic pattern that<br />
they inherited from their common ancestor.<br />
• Eyes have evolved perhaps 40 separate times in the evolutionary<br />
history <strong>of</strong> animals (see above). All <strong>of</strong> these animals<br />
share an organizer gene, called pax, that stimulates the<br />
development <strong>of</strong> eyes. In each <strong>of</strong> these lineages, the actual<br />
structure <strong>of</strong> the eye has evolved separately. There is a<br />
marine worm that has compound eyes, like those <strong>of</strong> other<br />
invertebrates, but also has brain lobes that have photosensitive<br />
chemicals similar to those <strong>of</strong> vertebrate eyes. Apparently<br />
vertebrate eyes evolved from invertebrate brain lobes<br />
rather than from invertebrate eyes.<br />
In modern organisms, scientists can recognize convergence,<br />
because despite the external similarities, the organisms<br />
are internally different and have achieved the convergence in<br />
different ways. In looking at fossils, scientists cannot always<br />
tell whether organisms shared a characteristic because they<br />
inherited it from a common ancestor, or because they evolved<br />
it separately by convergence. This sometimes makes reconstructing<br />
fossil history difficult. Sometimes in the fossil record<br />
a species appears to have become extinct then reappears. Is it<br />
because the species was rare for a long time, then reappeared<br />
in greater abundance (what paleontologist David Jablonski<br />
calls Lazarus taxa, because they appeared to rise from the<br />
dead), or because another lineage evolved similar features<br />
by convergence, and it just looked like a reappearance (what<br />
paleontologist Douglas Erwin calls Elvis taxa)?<br />
Every evolutionary scientist understands that there are<br />
many examples <strong>of</strong> convergent evolution. They differ in how<br />
much importance they give convergence in the overall pattern<br />
<strong>of</strong> the evolution <strong>of</strong> life. Some evolutionary scientists claim<br />
that evolution has so many different possible directions that<br />
the actual course taken by evolution is a matter <strong>of</strong> historical<br />
contingency, a viewpoint expressed by evolutionary biologist<br />
Stephen Jay Gould (see Gould, Stephen Jay; progress,<br />
concept <strong>of</strong>). Gould said that evolution produced a bush <strong>of</strong><br />
different forms, not a ladder leading upward to human perfection.<br />
These scientists consider convergence to be details<br />
that have occurred at the tips <strong>of</strong> the branches <strong>of</strong> the bush <strong>of</strong><br />
life. Others, like paleontologist Simon Conway Morris, insist<br />
that convergence is so frequent that it constitutes one <strong>of</strong> the<br />
major features <strong>of</strong> evolution. They point out that even though<br />
there may be an almost infinite number <strong>of</strong> possible biological<br />
adaptations, there are only a limited number <strong>of</strong> strategies<br />
that work, and evolution keeps finding these strategies over<br />
Copernicus, Nicolaus<br />
and over. Conway Morris agrees with Gould that evolution<br />
is a bush, not a ladder; but in this bush, he says, many <strong>of</strong> the<br />
branches are parallel. Conway Morris cites the parallel evolution<br />
<strong>of</strong> large brains in cetaceans (such as dolphins) and in<br />
humans. In the human lineage, there was a separate increase<br />
in brain size in the lineage that led to Homo sapiens, the lineage<br />
that led to Neandertals, perhaps also in the lineages<br />
that led to Asian Homo erectus; and in the robust australopithecines.<br />
Agriculture and civilization in humans evolved<br />
separately in East Asia, the Middle East, and America. Intelligence<br />
does not always <strong>of</strong>fer an evolutionary advantage (see<br />
intelligence, evolution <strong>of</strong>), but when it does, one would<br />
expect that the large processing center for the nerves would<br />
be near the front end <strong>of</strong> the animal, where most <strong>of</strong> the sensory<br />
organs are located (a big brain in a head with eyes and nose).<br />
Conway Morris says that the evolution <strong>of</strong> intelligence is a rare<br />
event, but that if humans encounter an intelligent life-form<br />
somewhere else in the universe it would probably be something<br />
that they would recognize as a person, perhaps even as a<br />
human. This is why he refers to “inevitable humans in a lonely<br />
universe” (see essay, “Are Humans Alone in the Universe?”).<br />
Further <strong>Reading</strong><br />
Conway Morris, Simon. Life’s Solution: Inevitable Humans in a<br />
Lonely Universe. Cambridge University Press, 2003.<br />
Mundy, N. I., et al. “Conserved genetic basis <strong>of</strong> a quantitative plumage<br />
trait involved in mate choice.” Science 303 (2004): 1,870–1,873.<br />
Summarized by Hoekstra, Hopi E., and Trevor Price. “Parallel evolution<br />
is in the genes.” Science 303 (2004): 1,779–1,781.<br />
Patterson, Thomas B., and Thomas J. Givnish. “Geographic cohesion,<br />
chromosomal evolution, parallel adaptive radiations, and consequent<br />
floral adaptations in Calochortus (Calochortaceae): Evidence<br />
from a cpDNA phylogeny.” New Phytologist 161 (2003): 253–264.<br />
Pichersky, Eran. “Plant scents.” American Scientist 92 (2004): 514–<br />
521.<br />
Shadwick, Robert E. “How tunas and lamnid sharks swim: An evolutionary<br />
convergence.” American Scientist 93 (2005): 524–531.<br />
Weinrech, Daniel M., et al. “Darwinian evolution can follow only<br />
very few mutational paths to fitter proteins.” Science 312 (2006):<br />
111–114.<br />
Copernicus, Nicolaus (1473–1543) Polish Astronomer<br />
Mikołaj Koppernigk (who Latinized his name to Nicolaus<br />
Copernicus) was born February 19, 1473. He attended university<br />
in Krakow, Poland, and appeared to be headed for<br />
a church career. But he also studied astronomy, geography,<br />
and mathematics and was particularly interested in planetary<br />
theory and patterns <strong>of</strong> eclipses. He was <strong>of</strong>fered a church<br />
administrative position but delayed taking it so that he could<br />
continue scientific studies in astronomy and medicine, which<br />
he undertook in Italy. He then returned to Poland and continued<br />
studying astronomy while he performed church duties.<br />
At the time, astronomers considered the Earth to be the<br />
center <strong>of</strong> the universe. The Sun and the planets circled the<br />
Earth. This belief made it difficult to explain the apparent<br />
movements <strong>of</strong> the planets in the sky, especially when the planets<br />
appeared to reverse their motion temporarily. Astronomers