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Encyclopedia of Evolution.pdf - Online Reading Center

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fossils and fossilization<br />

The grain <strong>of</strong> the wood is still visible in a petrified trunk <strong>of</strong> a tree that<br />

lived about 00 million years ago, now in Petrified Forest National Park,<br />

Arizona. (Photograph by Stanley A. Rice)<br />

derived. The chemical differences between the fossil and its<br />

matrix can sometimes be <strong>of</strong> help in extracting the fossil. For<br />

example, shells <strong>of</strong> invertebrates can sometimes be isolated if<br />

the matrix is dissolved in concentrated hydr<strong>of</strong>luoric acid.<br />

Sediments are the major, but not the only, place to find<br />

fossils. Many insects, stumbling into conifer sap, became<br />

entangled and engulfed; the sap became amber, and the<br />

insects were preserved in exquisite detail. This happened frequently<br />

enough that there is a worldwide market for insects<br />

preserved in amber.<br />

Ancient wood other than petrified wood, human bodies<br />

from a bog, and mummies preserved in Andean caves are not<br />

usually considered fossils, nor are bodies preserved by human<br />

activity (such as Egyptian mummies). Fossils retain the structure<br />

<strong>of</strong> the original organisms, sometimes with considerable<br />

internal detail; therefore coal and oil are not considered fossils,<br />

since they have been transformed into uniform organic<br />

materials (even though they are called fossil fuels). These<br />

remains can be as valuable as fossils in scientific research.<br />

Leaves that have been preserved for 20 million years in cold,<br />

wet, anoxic conditions in deposits near Clarkia, Idaho, are<br />

still green but oxidize into black films upon exposure to air.<br />

The leaves were so chemically intact that scientists could<br />

extract chloroplast DNA from them (see DNA [evidence<br />

for evolution]).<br />

The hard parts <strong>of</strong> organisms (such as the calcium carbonate<br />

shells <strong>of</strong> invertebrates, and the calcium phosphate bones<br />

<strong>of</strong> vertebrates) are most easily preserved and constitute the<br />

vast majority <strong>of</strong> fossils. Teeth are particularly popular fossils<br />

to study, not only because they are hard (and abundantly preserved)<br />

but also because they can reveal what kind <strong>of</strong> diet the<br />

animal had—grinding teeth for grazers, sharp teeth for carnivores.<br />

The study <strong>of</strong> fossil bones can also reveal more than<br />

just the anatomical structure <strong>of</strong> the animal. Study <strong>of</strong> tissue<br />

layers inside <strong>of</strong> bones reveals the growth rate <strong>of</strong> the animal<br />

and its age at death. This is how scientists have determined<br />

that Tyrannosaurus dinosaurs had a high metabolic rate and<br />

grew rapidly, up to 880 pounds (400 kg) by the age <strong>of</strong> 20<br />

years (see dinosaurs).<br />

Under conditions <strong>of</strong> very slow decomposition and<br />

very thorough burial, s<strong>of</strong>t parts <strong>of</strong> organisms, and entire<br />

organisms without hard parts, can be preserved. A famous<br />

example is the Burgess shale in Canada. Impressions <strong>of</strong><br />

s<strong>of</strong>t-bodied animals, many parts <strong>of</strong> plants, and even singlecelled<br />

microbes have frequently been preserved. Since fossilization<br />

almost always requires burial in sediment, most<br />

fossils are <strong>of</strong> the aquatic organisms that already live in or<br />

near the sediments. Aquatic invertebrates therefore constitute<br />

the best fossil assemblages. Terrestrial organisms are<br />

fossilized when they fall into or are washed into the sediments,<br />

as when primitive birds fell into low-oxygen pools<br />

that later became the limestone deposits <strong>of</strong> Solnh<strong>of</strong>en limestone<br />

in Germany (see arcHaeopteryx). The odds are<br />

very much against the preservation <strong>of</strong> fossils from terrestrial<br />

organisms: It is estimated that only one bone in a billion is<br />

preserved as a fossil. If all 290 million Americans, most <strong>of</strong><br />

whom have 206 bones, were to die, only about 60 bones<br />

would be fossilized under normal conditions. The odds<br />

are not even good for organisms <strong>of</strong> shallow oceans, where<br />

only 40 percent <strong>of</strong> the organisms have even the possibility<br />

<strong>of</strong> being fossilized. Rarely, massive graveyards <strong>of</strong> fossils<br />

(Lagerstätten) such as the Burgess shale or the Solnh<strong>of</strong>en<br />

limestone have resulted from conditions that were remarkably<br />

good for preservation.<br />

An intimate knowledge <strong>of</strong> biology is necessary in the<br />

study <strong>of</strong> fossils. This allows a great deal <strong>of</strong> information about<br />

the living animal to be reconstructed from the fossils <strong>of</strong> just<br />

its bones. Sites <strong>of</strong> muscle attachment leave traces on the<br />

bones, therefore the musclature <strong>of</strong> an extinct animal can be<br />

reconstructed from bone fossils. Blood vessels inside the living<br />

bones may have left traces in the fossilized bones, allowing<br />

scientists to determine the extent <strong>of</strong> blood vessel formation,<br />

and hence whether the animal was warm-blooded. Occasional<br />

lucky finds reveal even more about the animal’s life.<br />

Gastroliths are smooth stones found associated with dinosaur<br />

skeletons. Like modern birds (which are classified with<br />

the dinosaurs; see birds, evolution <strong>of</strong>), many dinosaurs<br />

swallowed stones which helped to grind plant food in their<br />

stomachs. The grinding action caused the stones to become<br />

smooth. Coprolites (fossilized feces) <strong>of</strong>ten allow scientists to<br />

reconstruct the diet <strong>of</strong> the animal.

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