Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
You also want an ePaper? Increase the reach of your titles
YUMPU automatically turns print PDFs into web optimized ePapers that Google loves.
fossils and fossilization<br />
The grain <strong>of</strong> the wood is still visible in a petrified trunk <strong>of</strong> a tree that<br />
lived about 00 million years ago, now in Petrified Forest National Park,<br />
Arizona. (Photograph by Stanley A. Rice)<br />
derived. The chemical differences between the fossil and its<br />
matrix can sometimes be <strong>of</strong> help in extracting the fossil. For<br />
example, shells <strong>of</strong> invertebrates can sometimes be isolated if<br />
the matrix is dissolved in concentrated hydr<strong>of</strong>luoric acid.<br />
Sediments are the major, but not the only, place to find<br />
fossils. Many insects, stumbling into conifer sap, became<br />
entangled and engulfed; the sap became amber, and the<br />
insects were preserved in exquisite detail. This happened frequently<br />
enough that there is a worldwide market for insects<br />
preserved in amber.<br />
Ancient wood other than petrified wood, human bodies<br />
from a bog, and mummies preserved in Andean caves are not<br />
usually considered fossils, nor are bodies preserved by human<br />
activity (such as Egyptian mummies). Fossils retain the structure<br />
<strong>of</strong> the original organisms, sometimes with considerable<br />
internal detail; therefore coal and oil are not considered fossils,<br />
since they have been transformed into uniform organic<br />
materials (even though they are called fossil fuels). These<br />
remains can be as valuable as fossils in scientific research.<br />
Leaves that have been preserved for 20 million years in cold,<br />
wet, anoxic conditions in deposits near Clarkia, Idaho, are<br />
still green but oxidize into black films upon exposure to air.<br />
The leaves were so chemically intact that scientists could<br />
extract chloroplast DNA from them (see DNA [evidence<br />
for evolution]).<br />
The hard parts <strong>of</strong> organisms (such as the calcium carbonate<br />
shells <strong>of</strong> invertebrates, and the calcium phosphate bones<br />
<strong>of</strong> vertebrates) are most easily preserved and constitute the<br />
vast majority <strong>of</strong> fossils. Teeth are particularly popular fossils<br />
to study, not only because they are hard (and abundantly preserved)<br />
but also because they can reveal what kind <strong>of</strong> diet the<br />
animal had—grinding teeth for grazers, sharp teeth for carnivores.<br />
The study <strong>of</strong> fossil bones can also reveal more than<br />
just the anatomical structure <strong>of</strong> the animal. Study <strong>of</strong> tissue<br />
layers inside <strong>of</strong> bones reveals the growth rate <strong>of</strong> the animal<br />
and its age at death. This is how scientists have determined<br />
that Tyrannosaurus dinosaurs had a high metabolic rate and<br />
grew rapidly, up to 880 pounds (400 kg) by the age <strong>of</strong> 20<br />
years (see dinosaurs).<br />
Under conditions <strong>of</strong> very slow decomposition and<br />
very thorough burial, s<strong>of</strong>t parts <strong>of</strong> organisms, and entire<br />
organisms without hard parts, can be preserved. A famous<br />
example is the Burgess shale in Canada. Impressions <strong>of</strong><br />
s<strong>of</strong>t-bodied animals, many parts <strong>of</strong> plants, and even singlecelled<br />
microbes have frequently been preserved. Since fossilization<br />
almost always requires burial in sediment, most<br />
fossils are <strong>of</strong> the aquatic organisms that already live in or<br />
near the sediments. Aquatic invertebrates therefore constitute<br />
the best fossil assemblages. Terrestrial organisms are<br />
fossilized when they fall into or are washed into the sediments,<br />
as when primitive birds fell into low-oxygen pools<br />
that later became the limestone deposits <strong>of</strong> Solnh<strong>of</strong>en limestone<br />
in Germany (see arcHaeopteryx). The odds are<br />
very much against the preservation <strong>of</strong> fossils from terrestrial<br />
organisms: It is estimated that only one bone in a billion is<br />
preserved as a fossil. If all 290 million Americans, most <strong>of</strong><br />
whom have 206 bones, were to die, only about 60 bones<br />
would be fossilized under normal conditions. The odds<br />
are not even good for organisms <strong>of</strong> shallow oceans, where<br />
only 40 percent <strong>of</strong> the organisms have even the possibility<br />
<strong>of</strong> being fossilized. Rarely, massive graveyards <strong>of</strong> fossils<br />
(Lagerstätten) such as the Burgess shale or the Solnh<strong>of</strong>en<br />
limestone have resulted from conditions that were remarkably<br />
good for preservation.<br />
An intimate knowledge <strong>of</strong> biology is necessary in the<br />
study <strong>of</strong> fossils. This allows a great deal <strong>of</strong> information about<br />
the living animal to be reconstructed from the fossils <strong>of</strong> just<br />
its bones. Sites <strong>of</strong> muscle attachment leave traces on the<br />
bones, therefore the musclature <strong>of</strong> an extinct animal can be<br />
reconstructed from bone fossils. Blood vessels inside the living<br />
bones may have left traces in the fossilized bones, allowing<br />
scientists to determine the extent <strong>of</strong> blood vessel formation,<br />
and hence whether the animal was warm-blooded. Occasional<br />
lucky finds reveal even more about the animal’s life.<br />
Gastroliths are smooth stones found associated with dinosaur<br />
skeletons. Like modern birds (which are classified with<br />
the dinosaurs; see birds, evolution <strong>of</strong>), many dinosaurs<br />
swallowed stones which helped to grind plant food in their<br />
stomachs. The grinding action caused the stones to become<br />
smooth. Coprolites (fossilized feces) <strong>of</strong>ten allow scientists to<br />
reconstruct the diet <strong>of</strong> the animal.