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Encyclopedia of Evolution.pdf - Online Reading Center

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A map <strong>of</strong> the Galápagos Islands, <strong>of</strong>f the coast <strong>of</strong> Ecuador<br />

Meskhize, Nicholas, and Athanasios Nenes. “Phytoplankton and<br />

cloudiness in the southern ocean.” Science 314 (2006): 1419–1423.<br />

Malin, G., and G. O. Kirst. “Algal production <strong>of</strong> dimethyl sulfide and<br />

its atmospheric role.” Journal <strong>of</strong> Phycology 33 (1997): 889–896.<br />

Margulis, Lynn, and Dorion Sagan. Slanted Truths: Essays on Gaia,<br />

Symbiosis, and <strong>Evolution</strong>. New York: Copernicus, 1997.<br />

Galápagos Islands The Galápagos Islands are volcanic<br />

islands that have played and continue to play a very important<br />

role in evolutionary science. They belong to Ecuador and<br />

are about 600 miles (1,000 km) <strong>of</strong>f the coast <strong>of</strong> that country<br />

(see figure above). Much <strong>of</strong> the volcanic activity that produced<br />

them occurred about five million years ago, which is<br />

very recent in evolutionary time.<br />

Being directly on the equator, these islands experience<br />

almost no variation in sunlight. They are right in the path <strong>of</strong><br />

oceanic currents that influence temperature and moisture conditions.<br />

There is an annual fluctuation <strong>of</strong> temperature caused<br />

by the ocean currents. Because <strong>of</strong> this, the cactuses can use<br />

a temperature cue that allows them to bloom every January.<br />

The El Niño Southern Oscillation alternates between dry<br />

conditions, when the ocean water near the South American<br />

coast is cool, and wet conditions, when the water is warm.<br />

During an El Niño year, large amounts <strong>of</strong> rainfall stimulate<br />

plant growth, particularly <strong>of</strong> grasses, vines, and weeds, but<br />

diminish the productivity <strong>of</strong> the oceans. The food chain <strong>of</strong> the<br />

ocean depends upon the minerals brought up from cool, deep<br />

ocean waters; when the water is warm, these minerals are not<br />

available. However, periods <strong>of</strong> heavy rainfall are uncommon,<br />

Galápagos Islands<br />

and the lower slopes <strong>of</strong> the Galápagos Islands are usually hot<br />

and dry, with scattered trees and a lot <strong>of</strong> hot, bare ground.<br />

The upland areas <strong>of</strong> the larger islands are frequently shrouded<br />

in mist, which provides enough moisture for forests to grow.<br />

The islands are famous for their unusual plant and animal<br />

life. There are no large native grazing animals or predators. The<br />

native animals have evolved, until recently, in the absence <strong>of</strong><br />

predators and human contact. From the fur seals to the groundnesting<br />

birds, they show no fear <strong>of</strong> humans. Many <strong>of</strong> the species<br />

are unique to these islands, or almost so. This is because<br />

they evolved on the islands from small founding populations<br />

<strong>of</strong> mainland species. All over the world, islands are home to<br />

unique species (see biogeography). The unique species found<br />

on islands was one <strong>of</strong> the observations that led Charles Darwin<br />

(see Darwin, Charles) to begin thinking about the process <strong>of</strong><br />

evolution. During his round-the-world voyage on HMS Beagle,<br />

he visited many islands, including the Galápagos. His 1835<br />

visit to the Galápagos turned out to be perhaps the single most<br />

important event that sparked Darwin’s understanding that evolution<br />

occurred by means <strong>of</strong> natural selection.<br />

Among the plants and animals that have undergone adaptive<br />

radiation on the Galápagos Islands are:<br />

• Finches. These birds have evolved from a single founding<br />

population into about 14 species that specialize on different<br />

ways <strong>of</strong> life and types <strong>of</strong> food (see Darwin’s finches).<br />

Ever since David Lack studied them in 1947, they have<br />

Iguanas bask in the sunlight on the stone marker <strong>of</strong> Ecuador’s Galápagos<br />

Islands National Park. These iguanas are just one <strong>of</strong> many species <strong>of</strong><br />

organisms unique to these islands. (Photograph by Stanley A. Rice)

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