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Encyclopedia of Evolution.pdf - Online Reading Center

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iology<br />

D. Response to environment. Both plants and animals<br />

have mechanisms that detect environmental stimuli, and processes<br />

that allow a response to them. In plants stimuli are usually<br />

detected by molecules such as pigments, and hormones<br />

allow the responses, such as bending toward light. Animals<br />

have sensory organs and a central nervous system that figures<br />

out the appropriate response to the stimuli.<br />

V. Sexual reproduction. Nearly all organisms have sexual life<br />

cycles. Certain organs produce cells with only half the genetic<br />

information that normal cells contain (see meiosis). In many<br />

plants and animals, these reproductive cells come in two sizes:<br />

The large female ones are either megaspores or eggs, while<br />

the small male ones are either microspores or sperm. The<br />

male cells <strong>of</strong> one individual may fuse with the female cells <strong>of</strong><br />

another, a process called fertilization. Fertilization produces a<br />

zygote, which may be sheltered inside <strong>of</strong> a seed, an eggshell,<br />

or a womb (see sex, evolution <strong>of</strong>; life history, evolution<br />

<strong>of</strong>). Sexual reproduction is not necessary for survival<br />

but generates genetic diversity.<br />

VI. Inheritance patterns. Because the traits <strong>of</strong> different<br />

parents can be shuffled into new combinations by sexual<br />

reproduction, new combinations <strong>of</strong> traits can occur in each<br />

generation (see Mendelian genetics).<br />

VII. Populations and evolution. All the interacting organisms<br />

<strong>of</strong> one species in one location constitute a population.<br />

Populations can grow rapidly, but limited resources prevent<br />

them from doing so forever. Because some individuals in a<br />

population reproduce more successfully than others, natural<br />

selection occurs and the population evolves (see natural<br />

selection). If the population separates into two populations,<br />

they can evolve into two species (see speciation).<br />

VIII. Diversity. <strong>Evolution</strong> has produced millions <strong>of</strong> species.<br />

Biologists have attempted to classify organisms on the basis<br />

<strong>of</strong> their evolutionary diversification rather than merely on the<br />

differences in their appearance. One such classification is:<br />

A. Domain Archaea: bacteria-like cells that, today, survive<br />

in extreme environments (see archaebacteria).<br />

B. Domain Eubacteria: bacteria-like cells that may use<br />

sunlight (photosynthetic bacteria) or inorganic chemicals<br />

(chemosynthetic bacteria) as energy sources for producing<br />

organic molecules or may obtain energy from organic molecules<br />

in living or dead organisms (heterotrophic bacteria)<br />

(see bacteria, evolution <strong>of</strong>). Early in evolutionary history,<br />

some heterotrophic bacteria invaded larger cells, and today<br />

they are the mitochondria that release energy from sugar in<br />

nearly all larger cells. Early in evolutionary history, some<br />

photosynthetic bacteria invaded larger cells, and today they<br />

are the chloroplasts that carry out photosynthesis in some<br />

larger cells (see symbiogenesis).<br />

C. Domain Eukarya: organisms composed <strong>of</strong> complex<br />

cells with DNA in nuclei (see eukaryotes, evolution <strong>of</strong>).<br />

The evolutionary linages <strong>of</strong> the eukaryotes are still being<br />

worked out. From within these lineages evolved:<br />

1. The plant kingdom. Land plants are descendants<br />

<strong>of</strong> green algae (see seedless plants, evolution <strong>of</strong>; gymnosperms,<br />

evolution <strong>of</strong>; angiosperms, evolution <strong>of</strong>).<br />

2. The fungus kingdom consists <strong>of</strong> decomposers and<br />

pathogens that absorb food molecules.<br />

3. The animal kingdom. Animals, descendants <strong>of</strong> one<br />

<strong>of</strong> the lineages <strong>of</strong> protozoa (see invertebrates, evolution<br />

<strong>of</strong>; fishes, evolution <strong>of</strong>; amphibians, evolution <strong>of</strong>; reptiles,<br />

evolution <strong>of</strong>; birds, evolution <strong>of</strong>; mammals, evolution<br />

<strong>of</strong>).<br />

IX. Synecological context. This is the ecological interaction<br />

<strong>of</strong> organisms with one another (syn- comes from the Greek<br />

for together).<br />

A. General ecological interactions. <strong>Evolution</strong> has refined<br />

the interactions that broad groups <strong>of</strong> organisms have with<br />

one another (see coevolution): for example, herbivores that<br />

eat plants and plants that defend themselves from herbivores;<br />

predators that eat prey, and prey that defend themselves<br />

from predators; pollinators and flowering plants. Many complex<br />

animal behavior patterns have evolved, particularly in<br />

response to sexual selection.<br />

B. Symbiotic ecological interactions. Symbiosis results<br />

from the very close interaction <strong>of</strong> two species, in which at<br />

least one <strong>of</strong> the species depends upon the other. These interactions<br />

have resulted from coevolution.<br />

1. Parasitism occurs when a parasite harms the host.<br />

2. Commensalism occurs when a commensal has no<br />

effect on the host.<br />

3. Mutualism occurs when both species benefit.<br />

Mutualism is so widespread that, in some cases, different species<br />

<strong>of</strong> organisms have actually fused together and formed<br />

new kinds <strong>of</strong> organisms.<br />

4. Natural selection favors the evolution <strong>of</strong> hosts<br />

that resist parasites and sometimes favors the evolution <strong>of</strong><br />

parasites that have only mild effects on their hosts. Natural<br />

selection can favor the evolution <strong>of</strong> parasitism into commensalism,<br />

and commensalism into mutualism.<br />

C. Ecological communities are all <strong>of</strong> the interacting species<br />

in a location. They generally form clusters, based upon<br />

temperature and moisture conditions in different parts <strong>of</strong> the<br />

world: for example, tundra, forests, grasslands, deserts, lakes,<br />

shallow seas, deep oceans. Each community contains microhabitats<br />

with smaller communities within them.<br />

D. Ecological communities continually undergo change.<br />

On a large scale, continents drift (see continental drift),<br />

mass extinctions occur, and climates fluctuate (see ice<br />

ages; Snowball Earth). On a small scale, disturbances<br />

such as fires are followed by stages <strong>of</strong> regrowth called<br />

ecological succession. Billions <strong>of</strong> years <strong>of</strong> these changes<br />

have produced an entire evolutionary history, as described<br />

throughout this book, and the world as humans know it<br />

today.<br />

The Earth is filled with living organisms. But is the Earth<br />

itself alive? It does not match all <strong>of</strong> the characteristics listed<br />

above to qualify as an organism, but it does have some processes<br />

that produce a semblance <strong>of</strong> homeostasis, which is not<br />

readily understandable by the operations <strong>of</strong> the organisms on

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