Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
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dinosaurs<br />
what DeVries thought were mutations were actually recombinations<br />
<strong>of</strong> existing genes.<br />
DeVries was one <strong>of</strong> three biologists (along with Erich<br />
von Tschermak-Seysenegg and Carl Correns) who rediscovered<br />
the work <strong>of</strong> Gregor Mendel (see Mendel, Gregor;<br />
Mendelian genetics). Mendel had demonstrated that traits<br />
were passed on from one generation to another in a particulate<br />
fashion rather than by blending, and DeVries believed<br />
it was pangenes that did this. Largely because <strong>of</strong> DeVries,<br />
who influenced later geneticists such as Henry Bateson and<br />
Thomas Hunt Morgan, Mendel’s mutations were considered<br />
inconsistent with Darwinian evolution. It was not until the<br />
modern synthesis <strong>of</strong> the late 1930s that Darwinian evolution<br />
and Mendelian genetics were brought together in a way<br />
that showed DeVries to have been wrong. Interestingly, this<br />
might not have happened had DeVries not helped to rediscover<br />
and publicize the work <strong>of</strong> Gregor Mendel. The rediscovery<br />
<strong>of</strong> Mendel’s work remains DeVries’s major impact on<br />
evolutionary science. He died May 21, 1935.<br />
dinosaurs The term dinosaur usually refers to reptiles,<br />
many <strong>of</strong> them large, that dominated the Earth during the<br />
Mesozoic era. The earliest dinosaurs lived during the Triassic<br />
period. Dinosaurs diversified into numerous forms<br />
during the Jurassic period and Cretaceous period, perhaps<br />
because one <strong>of</strong> the mass extinctions, which occurred<br />
at the end <strong>of</strong> the Triassic period, removed competition from<br />
other vertebrate groups. The dinosaurs themselves became<br />
extinct at the end <strong>of</strong> the Cretaceous period (see Cretaceous<br />
extinction).<br />
Modern phylogeny (see cladistics) classifies organisms<br />
based on evolutionary divergence. Since birds and mammals<br />
diverged from the group <strong>of</strong> vertebrates usually called<br />
reptiles, there is no such thing as reptiles unless one includes<br />
birds and mammals in the group (see reptiles, evolution<br />
<strong>of</strong>). Dinosaurs had many characteristics <strong>of</strong> skin, teeth, and<br />
bones that would remind observers <strong>of</strong> the reptiles with which<br />
they are familiar. In fact, since birds evolved from dinosaurs,<br />
the cladistic approach would indicate that the dinosaurs still<br />
exist: They are birds. Therefore the dinosaurs that most people<br />
think <strong>of</strong> are more properly called non-avian dinosaurs.<br />
Although dinosaurs are truly ancient, the therapsid (mammallike)<br />
reptile lineage had diverged before the dinosaur lineage,<br />
and true mammals already existed by the time the first dinosaurs<br />
roamed the Earth.<br />
The first dinosaurs to be studied were large, which is<br />
why Sir Richard Owen (see Owen, Richard) gave them the<br />
name dinosaurs (Greek for “terrible lizards”) in 1842. Many<br />
famous dinosaur discoveries came from the northern plains<br />
<strong>of</strong> the United States. In the late 19th century, almost 150 new<br />
dinosaur species were revealed by the excavations and reconstructions<br />
<strong>of</strong> American paleontologists Edward Drinker Cope<br />
and Othniel Charles Marsh. They began as collaborators but<br />
ended up in a three-decade intense rivalry. Marsh did not perform<br />
very much fieldwork. One account claims that he visited<br />
a spot where dinosaur bones were lying about like logs but<br />
did not recognize them. Marsh was, however, rich enough to<br />
buy fossils and to hire field workers. His uncle, George Peabody,<br />
endowed a museum at Yale specifically so that Marsh<br />
could study dinosaurs. Cope, who was also wealthy, spent<br />
much productive time in the field. He worked in Montana at<br />
the same time that General Custer’s army was being annihilated<br />
nearby. When suspicious Natives checked out his camp,<br />
he entertained them by taking out his false teeth until they<br />
went away. A schoolteacher who found some dinosaur bones<br />
genially informed both Marsh and Cope about them. Cope<br />
sent the schoolteacher money and told him to ask Marsh to<br />
send all the specimens back to him. The rivalry grew intense<br />
enough that the excavators <strong>of</strong> the Marsh and Cope camps<br />
threw rocks at one another. Cope lost his money, but not his<br />
ego, in financial speculation, dying poor. He willed his body<br />
to a museum, hoping to have his skeleton used as the “type<br />
specimen” to represent the human species. When the skeleton<br />
was prepared, it was found to have syphilitic lesions and was<br />
unsuitable as a type specimen.<br />
Most <strong>of</strong> what scientists know about dinosaurs comes<br />
from the study <strong>of</strong> fossilized bones, in the tradition <strong>of</strong> Cope<br />
and Marsh. Recently this has included the microscopic study<br />
<strong>of</strong> the spaces left by blood vessels in the bones. Other kinds<br />
<strong>of</strong> fossil evidence have also allowed many insights into the<br />
lives <strong>of</strong> dinosaurs:<br />
• Gastroliths. These are “stomach stones” that helped to<br />
grind coarse plant material in the stomachs <strong>of</strong> some dinosaurs<br />
that ate branches and leaves. Many modern birds<br />
swallow stones that help them to grind their food.<br />
• Eggs. Fossilized eggs with embryos inside reveal some <strong>of</strong> the<br />
process <strong>of</strong> dinosaur development. In addition, the discovery<br />
<strong>of</strong> caches <strong>of</strong> eggs reveals that dinosaurs cared for nests <strong>of</strong><br />
eggs in a manner similar to that <strong>of</strong> most modern birds.<br />
• S<strong>of</strong>t material. In 2005 paleontologist Mary Higby Schweitzer<br />
discovered preserved s<strong>of</strong>t tissue, including what<br />
may be blood vessels and cells, in a dinosaur bone.<br />
Since the time <strong>of</strong> Owen, Marsh, and Cope, dinosaurs<br />
have entered the popular imagination as emblematic both <strong>of</strong><br />
huge size and <strong>of</strong> evolutionary failure. In both ways, the image<br />
is unfair. First, many dinosaurs were small or medium sized.<br />
Many <strong>of</strong> the smaller dinosaurs were sleek and rapid, and<br />
some had feathers (see birds, evolution <strong>of</strong>). Second, there<br />
were many kinds <strong>of</strong> dinosaurs, all over the world; they were,<br />
for more than a hundred million years, an astounding evolutionary<br />
success.<br />
Some dinosaurs were extremely large. Sauroposeidon<br />
proteles, which lived 110 million years ago in what is now<br />
Oklahoma, had a body 100 feet (30 m) in length, with a<br />
neck more than 30 feet (10 m) long. The size estimate <strong>of</strong> the<br />
neck <strong>of</strong> this dinosaur is based upon the fact that most vertebrates<br />
have the same number <strong>of</strong> neck vertebrae, and the<br />
longest vertebrae found for this dinosaur were about five feet<br />
(140 cm) long. The sauropod Supersaurus weighed nearly<br />
a hundred tons, the largest animal ever to live on land (see<br />
figure on page 123). They were also the largest four-legged<br />
animals that would possibly have lived; if they had grown to<br />
140 tons, the size <strong>of</strong> the legs necessary to support the weight