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Encyclopedia of Evolution.pdf - Online Reading Center

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0 Cenozoic era<br />

opens the way to an understanding <strong>of</strong> mass extinctions,<br />

followed by rapid evolution, that punctuate the history <strong>of</strong> life<br />

on Earth (see punctuated equilibria). Modern geology, by<br />

presenting a background <strong>of</strong> uniformitarian processes punctuated<br />

by catastrophes such as asteroid impacts (see Permian<br />

extinction; Cretaceous extinction), has preserved the<br />

best <strong>of</strong> both 19th-century approaches: uniformitarianism and<br />

catastrophism.<br />

Further <strong>Reading</strong><br />

Benton, Michael. “The death <strong>of</strong> catastrophism.” Chap. 3 in When<br />

Life Nearly Died: The Greatest Mass Extinction <strong>of</strong> All Time.<br />

London: Thames and Hudson, 2003.<br />

Gould, Stephen Jay. “Uniformity and catastrophe.” Chap. 18 in Ever<br />

Since Darwin: Reflections in Natural History. New York: Norton,<br />

1977.<br />

Cenozoic era The Cenozoic era (the era <strong>of</strong> “recent life”)<br />

is the third era <strong>of</strong> the Phanerozoic Eon, or period <strong>of</strong> visible<br />

multicellular life, which followed the Precambrian time in<br />

Earth history (see geological time scale). The Cenozoic<br />

era, which is the current era <strong>of</strong> Earth history, began with the<br />

mass extinction event that occurred at the end <strong>of</strong> the Cretaceous<br />

period <strong>of</strong> the Mesozoic era (see mass extinctions;<br />

Cretaceous extinction). The Cenozoic era is traditionally<br />

divided into two geological periods, the Tertiary period<br />

and the Quaternary period.<br />

The Cretaceous extinction left a world in which many<br />

organisms had died and much space and many resources were<br />

available for growth. The dinosaurs had become extinct,<br />

as well as numerous lineages within the birds, reptiles, and<br />

mammals (see birds, evolution <strong>of</strong>; reptiles, evolution<br />

<strong>of</strong>; mammals, evolution <strong>of</strong>). The conifers that had dominated<br />

the early Mesozoic forests came to dominate only the<br />

forests <strong>of</strong> cold or nutrient-poor mountainous regions in the<br />

Cenozoic (see gymnosperms, evolution <strong>of</strong>). The flowering<br />

plants evolved in the Cretaceous period but proliferated during<br />

the Cenozoic era, into the forest trees that dominate the<br />

temperate and tropical regions, and many shrubs and herbaceous<br />

species (see angiosperms, evolution <strong>of</strong>). The explosive<br />

speciation <strong>of</strong> flowering plants paralleled that <strong>of</strong> insect<br />

groups such as bees, butterflies, and flies, a pattern most scientists<br />

attribute to coevolution.<br />

The Cenozoic world was cooler and drier than most <strong>of</strong><br />

previous Earth history. Climatic conditions became cooler and<br />

drier during the late Cenozoic than they had been early in the<br />

Cenozoic. During the middle <strong>of</strong> the Tertiary period, dry conditions<br />

allowed the adaptation <strong>of</strong> grasses and other plants to aridity,<br />

and the spread <strong>of</strong> grasslands and deserts. Grazing animals<br />

evolved from browsing ancestors, taking advantage <strong>of</strong> the grass<br />

food base (see horses, evolution <strong>of</strong>). Five million years ago,<br />

the Mediterranean dried up into a salt flat. The coolest and driest<br />

conditions began with the Quaternary period. About every<br />

hundred thousand years, glaciers build up around the Arctic<br />

Ocean and push southward over the northern continents (see<br />

ice ages). Meanwhile, lower sea levels and reduced evaporation<br />

result in drier conditions in the equatorial regions.<br />

Many evolutionary scientists and geologists now divide<br />

the Cenozoic era into the Paleogene (Paleocene, Eocene, Oligocene)<br />

and the Neogene (Miocene, Pliocene, Pleistocene,<br />

Holocene) rather than the traditional Tertiary and Quaternary<br />

periods.<br />

Chambers, Robert (1802–1871) British Publisher Born<br />

July 10, 1802, Robert Chambers was a British publisher,<br />

whose role in the development <strong>of</strong> evolutionary theory was<br />

not widely known until after his death. Long before Darwin<br />

(see Darwin, Charles) wrote his famous book (see origin<br />

<strong>of</strong> species [book]), evolutionary thought was in the air in<br />

Europe. The naturalist Buffon (see Buffon, Georges) had<br />

presented a limited evolutionary theory in France in the 18th<br />

century. Charles Darwin’s grandfather (see Darwin, Erasmus)<br />

had speculated about the possibility, and the French scientist<br />

Lamarck had proposed a scientific evolutionary theory<br />

(see Lamarckism). In October <strong>of</strong> 1844, an anonymous British<br />

book, Vestiges <strong>of</strong> the Natural History <strong>of</strong> Creation, presented<br />

an evolutionary history <strong>of</strong> the Earth, from the formation <strong>of</strong><br />

the solar system and plant and animal life, including even<br />

the origins <strong>of</strong> humankind. More than 20,000 copies sold in<br />

a decade. Political leaders like American president Abraham<br />

Lincoln, British Queen Victoria, and British statesmen William<br />

Gladstone and Benjamin Disraeli read it. Poets, such as Alfred<br />

Tennyson, and philosophers like John Stuart Mill did also.<br />

Many scientists also read it (see Huxley, Thomas Henry).<br />

The purpose <strong>of</strong> the book, said its author, was to provoke scientific<br />

and popular discussion about evolution.<br />

It certainly succeeded in this objective. Responses ranged<br />

from enthusiasm to condemnation. A British medical journal,<br />

Lancet, described Vestiges as “a breath <strong>of</strong> fresh air.” Physicist<br />

Sir David Brewster wrote that Vestiges raised the risk <strong>of</strong> “poisoning<br />

the fountains <strong>of</strong> science, and sapping the foundations<br />

<strong>of</strong> religion.” Scottish geologist Hugh Miller published an<br />

entire book, Foot-Prints <strong>of</strong> the Creator, as a rebuttal to Vestiges.<br />

Charles Darwin called it a “strange, unphilosophical,<br />

but capitally-written book,” and noted that some people had<br />

suspected him <strong>of</strong> being the author. Huxley recognized it as<br />

the work <strong>of</strong> an amateur whose author could “indulge in science<br />

at second-hand and dispense totally with logic.” A pr<strong>of</strong>essional<br />

scientist would have dismissed the fraudulent claim<br />

<strong>of</strong> the amateur scientist, Mr. W. H. Weekes, who claimed to<br />

have created living mites by passing electric currents through<br />

a solution <strong>of</strong> potassium ferrocyanate, a claim that the Vestiges<br />

author was credulous enough to believe.<br />

It was not until 1884, after Darwin’s death and evolutionary<br />

science had become respectable, that the author was<br />

<strong>of</strong>ficially revealed to be Robert Chambers, one <strong>of</strong> Britain’s<br />

most successful publishers. Chambers had chosen to remain<br />

anonymous because he feared the reputation <strong>of</strong> Vestiges<br />

would hurt his business. (His company published, among<br />

many other things, Bibles.) Because <strong>of</strong> Chambers’s interest in<br />

science, some people, including Darwin, had already guessed<br />

the identity <strong>of</strong> the notorious “Mr. Vestiges.”<br />

Chambers and his older brother had begun their publishing<br />

business by selling cheap Bibles and schoolbooks from

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