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Encyclopedia of Evolution.pdf - Online Reading Center

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Haeckel, Ernst (1834–1919) German <strong>Evolution</strong>ary biologist<br />

Ernst Heinrich Philipp August Haeckel is best known as<br />

the principal champion for Darwinian evolution in late 19thcentury<br />

continental Europe, and for evolutionary concepts<br />

that turned out to be wrong in their original form.<br />

Born February 16, 1834, Ernst Haeckel was a physician<br />

but in 1859, upon reading Darwin’s book (see Darwin,<br />

Charles; origin <strong>of</strong> species [book]), he abandoned his practice.<br />

He studied zoology at the University <strong>of</strong> Jena, then became<br />

a pr<strong>of</strong>essor <strong>of</strong> comparative anatomy at that university in 1862.<br />

He extensively studied the embryology <strong>of</strong> invertebrates (see<br />

invertebrates, evolution <strong>of</strong>) and a group <strong>of</strong> protists (see<br />

eukaryotes, evolution <strong>of</strong>) known as radiolarians.<br />

Although Haeckel defended evolution, he like many<br />

other prominent scientists did not agree with Darwin that<br />

natural selection was the mechanism <strong>of</strong> evolution. In<br />

particular, Haeckel and many other scientists accepted a version<br />

<strong>of</strong> Lamarckism. He also believed that evolution had a<br />

predetermined direction. This direction was not impelled by<br />

spiritual forces—as many other German scholars believed,<br />

following the tradition <strong>of</strong> the scientist and writer Johann<br />

Wolfgang von Goethe—but by natural law. This direction<br />

could be glimpsed in the stages <strong>of</strong> development <strong>of</strong> an animal’s<br />

embryo. As Haeckel described it in a still famous quote,<br />

“Ontogeny recapitulates phylogeny.” This means that an animal’s<br />

embryo, as it develops, passes through the same evolutionary<br />

stages by which its species evolved. Thus the human<br />

embryo goes through fish, amphibian, reptile, and earlier<br />

mammalian stages, according to Haeckel’s interpretation.<br />

While evolutionary biologists still accept this overall pattern,<br />

it is considered an effect rather than a cause, and they do not<br />

accept it as a natural law (see recapitulation). The figures<br />

that Haeckel published, illustrating stages <strong>of</strong> embryonic<br />

development, contained errors. In particular, Haeckel used<br />

dog embryos to fill in some missing stages in human embryonic<br />

development. The differences were minor and did not<br />

alter the overall pattern but tarnished Haeckel’s reputation<br />

H<br />

among evolutionary scientists. He remained popular with the<br />

reading public.<br />

Haeckel also championed eugenics. He stated that “Politics<br />

is applied biology,” by which he meant that evolution<br />

led upward toward white Europeans and that politics should<br />

advance the social and biological interests <strong>of</strong> white Europeans.<br />

After Haeckel’s death, the Nazis used this quote and his<br />

justifications for racism, nationalism, and social Darwinism<br />

to support their cause.<br />

Haeckel applied his scientific concepts broadly, as an<br />

amateur scholar in anthropology, psychology, and cosmology.<br />

He used evolution as the basis for attacking religious<br />

views. He could be considered the German counterpart <strong>of</strong> T.<br />

H. Huxley (see Huxley, Thomas Henry) but without Huxley’s<br />

rigorous regard for facts. Haeckel died August 8, 1919.<br />

Further <strong>Reading</strong><br />

Gasman, D. The Scientific Origins <strong>of</strong> National Socialism: Social Darwinism<br />

in Ernst Haeckel and the German Monist League. London:<br />

MacDonald, 1972.<br />

Museum <strong>of</strong> Paleontology, University <strong>of</strong> California, Berkeley. “Ernst<br />

Haeckel (1834–1919).” Available online. URL: http://www.ucmp.<br />

berkeley.edu/history/haeckel.html. Accessed April 28, 2005.<br />

Haldane, J. B. S. (1892–1964) British Zoologist John<br />

Burdon Sanderson Haldane was an idiosyncratic genius who<br />

contributed to the modern understanding <strong>of</strong> evolution in several<br />

important ways. His father, John Scott Haldane, was a<br />

pr<strong>of</strong>essor <strong>of</strong> physiology at Oxford University, whose diverse<br />

studies included parasites, heatstroke, and altitude sickness. J.<br />

S. Haldane also studied the intervals that were necessary during<br />

an ascent from deep diving in order to avoid the bends. A<br />

fearless researcher, J. S. Haldane even took notes on his symptoms<br />

while mildly poisoning himself with carbon monoxide.<br />

Born November 5, 1892, J. B. S. Haldane worked alongside<br />

his father, J. S. Haldane, and conducted his own experiments,<br />

even in childhood. As a teenager, he bred guinea pigs

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