Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
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0 Smith, William<br />
———. Tempo and Mode in <strong>Evolution</strong>. New York: Columbia University<br />
Press, 1944.<br />
———. This View <strong>of</strong> Life: The World <strong>of</strong> an <strong>Evolution</strong>ist. New York:<br />
Harcourt, Brace and World, 1964.<br />
Smith, William (1769–1839) British Geologist, Engineer William<br />
Smith was the geologist who figured out that fossils could<br />
be used to correlate sedimentary rock layers (see fossils and<br />
fossilization). If a sedimentary layer in one location contained<br />
an assemblage <strong>of</strong> fossils, and a sedimentary layer in another<br />
location contained the same or similar assemblage <strong>of</strong> fossils, the<br />
layers in both locations (even if separated by hundreds <strong>of</strong> miles)<br />
were almost certainly formed at the same time. In this way, the<br />
relative ages <strong>of</strong> fossil deposits can be determined, although their<br />
absolute ages cannot. Smith’s insight allowed geologists in the<br />
19th century to reconstruct the geological record <strong>of</strong> the Earth’s<br />
entire history, although they had no idea <strong>of</strong> how many millions<br />
<strong>of</strong> years the geological record spanned (see age <strong>of</strong> Earth;<br />
radiometric dating).<br />
Born March 23, 1769, William Smith received little formal<br />
education but spent much time collecting fossils as he<br />
grew up. He studied mapping and became a surveyor. Excelling<br />
in this skill, Smith spent six years supervising the digging<br />
<strong>of</strong> the Somerset Canal in southwestern England. Smith<br />
noticed that the fossils within a vertical section <strong>of</strong> sedimentary<br />
rocks were always in the same order from the bottom<br />
to the top <strong>of</strong> the section, and the sedimentary rock sections<br />
on one side <strong>of</strong> England matched those on the other side. Not<br />
every species was useful for correlating the strata <strong>of</strong> different<br />
locations; species that had very widespread distributions<br />
or that persisted for a long time were not suitable for precise<br />
correlation.<br />
Smith was not the first to construct geological maps but<br />
was the first to do so by fossils and not by the mineral composition<br />
<strong>of</strong> the rocks. When Smith finally secured money from<br />
private investors in 1812, he began a fossil-based geological<br />
map <strong>of</strong> all <strong>of</strong> England and Wales, a task he finished in 1815.<br />
Even though this map was initially disregarded by scientists,<br />
it is difficult to overestimate its importance. It was an essential<br />
base <strong>of</strong> information that allowed uniformitarianism to<br />
replace catastrophism and that allowed the insights <strong>of</strong> later<br />
geologists (see Lyell, Charles) and, eventually, evolutionary<br />
scientists (see Darwin, Charles). Smith died August 28,<br />
1839.<br />
Further <strong>Reading</strong><br />
Winchester, Simon, and Soun Vannithone. The Map That Changed<br />
the World: William Smith and the Birth <strong>of</strong> Modern Geology. New<br />
York: Perennial, 2002.<br />
Snowball Earth Periods <strong>of</strong> ice formation, global in extent,<br />
occurred during Precambrian time. The Sturtian Glaciation<br />
ended about 700 million years ago, the Marinoan Glaciation<br />
about 635 million years ago, and the Gaskiers Glaciation<br />
about 580 million years ago. There was an earlier period <strong>of</strong><br />
ice formation, about 2.4 billion years ago, about which much<br />
less is known, because there are far fewer deposits that have<br />
survived from that distant time. The ice formed on both land<br />
and sea, not only in polar and temperate latitudes but even<br />
in the tropics, which would have made the Earth look very<br />
much like a snowball. Because there may have been little<br />
open ocean, there would have been very little evaporation,<br />
hence very little rain or snow, once the snowball conditions<br />
formed.<br />
Calculations by Russian scientist Mikhail Budyko in the<br />
1960s suggested that the Earth could never have had such a<br />
period <strong>of</strong> cold temperatures, because if this had occurred,<br />
the Earth would have remained frozen forever. This would<br />
occur because <strong>of</strong> the albedo effect: The sunlight would<br />
have reflected from the white snow and ice, directly back<br />
into outer space, without being absorbed; the only sunlight<br />
that creates warmth is the sunlight that is absorbed. (This<br />
is why a stick lying on top <strong>of</strong> the snow on a sunny day will<br />
melt its way into the snow; the stick absorbs sunlight and<br />
becomes warm, while the snow does not.) Budyko’s calculations<br />
did not include the effects <strong>of</strong> volcanic eruptions.<br />
Eruptions would have created numerous unfrozen spots in<br />
the ocean and would have released carbon dioxide into the<br />
atmosphere. Carbon dioxide contributes to the greenhouse<br />
effect by absorbing infrared photons that are emitted by<br />
the Earth. Very little infrared light would have been emitted<br />
by a Snowball Earth, but as carbon dioxide levels built up<br />
over millions <strong>of</strong> years, enough heat might have accumulated<br />
to begin melting the ice. Two major processes by which carbon<br />
dioxide is removed from the air are the photosynthesis<br />
<strong>of</strong> organisms and the weathering <strong>of</strong> rocks. With so few<br />
organisms, and with the rocks and oceans covered by ice,<br />
there would have been almost nothing to remove carbon<br />
dioxide from the air, thus allowing it to accumulate. These<br />
processes would have allowed the Earth to emerge from its<br />
snowball condition.<br />
Snowball Earth is an example <strong>of</strong> a truly radical theory<br />
that has become widely accepted, in various forms, by the<br />
scientific community. Its principal proponents are geologists<br />
Paul H<strong>of</strong>fman and Daniel Schrag <strong>of</strong> Harvard University, basing<br />
their studies on earlier work by geologists Joseph Kirschvink<br />
and Brian Harland.<br />
• One line <strong>of</strong> evidence suggesting that there was a Snowball<br />
Earth is the massive accumulation <strong>of</strong> tillite dropstones in<br />
late Precambrian deposits. Dropstones are a diverse mixture<br />
<strong>of</strong> rocks that have been transported far from their<br />
places <strong>of</strong> origin and dropped onto the bottom <strong>of</strong> the ocean.<br />
It is generally agreed that only icebergs can carry dropstones<br />
for such distances. The large deposits <strong>of</strong> dropstones<br />
indicate that there must have been a lot <strong>of</strong> icebergs in the<br />
late Precambrian. These deposits are found from all over<br />
the world: Spitsbergen, China, Australia, Russia, Norway,<br />
Namibia, Newfoundland, and the Rocky Mountains.<br />
Therefore, the glaciation was global.<br />
• Another line <strong>of</strong> evidence is the presence <strong>of</strong> late Precambrian<br />
ironstones. When oxygen first began to be produced<br />
by photosynthesis <strong>of</strong> microscopic organisms, it reacted<br />
with iron in the ocean water and precipitated to the bottom<br />
<strong>of</strong> the sea, forming iron deposits. This stopped happening<br />
when the iron was removed from the ocean water,